Railway flora

 

 

Vegetation has conquered this abandoned railway near the small town of San Cataldo, Sicily. (Photo copyright © by Kaj Halberg)

 

 

 

Vegetation has invaded this abandoned railway tracks in Nyborg, Denmark, including false oat-grass (Arrhenatherum elatius), red fescue (Festuca rubra), wild chervil (Anthriscus sylvestris), common mugwort (Artemisia vulgaris), dovesfoot crane’s-bill (Geranium molle), common goat’s-beard (Tragopogon pratensis), dandelion (Taraxacum officinale), narrow-leaved plantain (Plantago lanceolata), a species of burdock (Arctium), white clover (Trifolium repens), and traveller’s joy (Clematis vitalba). (Foto copyright © by Kaj Halberg)

 

 

 

Originally, the Jiancing Line Lumber Transportation Track, eastern Taiwan, was about 5.5 km long. As its name implies, it was constructed to bring out timber from the Taipingshan Mountains, part of which today constitutes the Taipingshan National Forest Recreation Area.

Jiancing Historic Trail was established along remains of this old railroad track. Besides the tracks themselves, you may study skids, switches, trolley axles, and deserted workers’ dwellings.

Jiancing means ‘seeing the sun’. The name was possibly given due to the fact that this area is often enveloped in fog, and finally seeing the sun breaking through the fog might be relieving.

 

 

Abandoned railroad tracks, overgrown by mosses, ferns, and other plants, Jiancing Historic Trail, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Many plants have adapted to a life along railways, which often constitute a refuge – at least until the worker arrives with chemical herbicides or a mower.

 

 

Amaranthaceae Amaranth family

 

Amaranthus Pigweed, amaranth
A worldwide genus with about 75 species.

In his delightful book All about Weeds, Dover Publications (1974), American botanist Edwin Spencer (1881-1964) says about a species of pigweed, A. retroflexus:“One of the most robust, devil-may-care weeds is the pigweed. ‘Careless weed’ is another of its common names, and it comes by this name honestly enough. In good rich soil, it cares for nothing. Wind, hail, fair weather and foul are all the same to the pigweed. Nor does it care what plants are its competitors. It can usually shoulder our any plant within reach, and it has a considerable reach. The name pigweed, however, has no reference to the piggish nature of the plant. It refers to the gustatory pleasure the weed affords pigs. Hogs will leave their corn to feast on pigweeds. In spite of its bristly appearance (it always reminds one of a boisterous young sailor with a week’s growth of sandy beard on his face and his cap on the side of his head), the leaves are tender, and if the smacking of lips and satisfied grunts mean the same thing to pigs that they do to Man, the weeds must be delicious.”

The generic name is derived from Ancient Greek a (‘not’, ‘without’) and maraino (‘to wither’), given in reference to the flowers, which last a very long time.

 

Amaranthus viridis Slender amaranth
The native area of this species, also known as green amaranth, is unknown, and today it is distributed in all warmer areas of the world.

Leaves, as well as seeds, are eaten in many parts of the world. In Australia, this plant was also a source of food during the 19th Century. In 1889, botanist Joseph Maiden (1859-1925) wrote: “It is an excellent substitute for spinach, being far superior to much of the leaves of the white beet sold for spinach in Sydney. Next to spinach it seems to be most like boiled nettle leaves, which when young are used in England, and are excellent. This amarantus should be cooked like spinach, and as it becomes more widely known, it is sure to be popular, except amongst persons who may consider it beneath their dignity to have anything to do with so common a weed.” (Source: T. Low, 1985. Wild Herbs of Australia & New Zealand. Angus & Robertson)

In India, slender amaranth is utilized in traditional Ayurvedic medicine.

The specific name is Latin, meaning ‘green’.

 

 

Prostrate form of slender amaranth, growing at a fence, Long Bien Railway Station, Hanoi, Vietnam. (Photo copyright © by Kaj Halberg)

 

 

 

Apiaceae Umbellifers

 

Ferula
A large genus with about 220 species, found from southern Europe and southern Russia southwards to northern and eastern Africa, Arabia, Tibet, and southern China, mostly growing in arid areas.

The generic name is Latin, meaning ‘rod’ or ‘staff’. In Ancient Greek mythology, Prometheus, the god of fire, presented fire to humans by hiding it in the hollow stem of giant fennel. The stem, entwined with foliage of ivy (Hedera helix), was also used as a thyrsus – a staff associated with the wine god Dionysus.

 

Ferula communis Giant fennel
This huge plant is distributed around the Mediterranean, on the Arabian Peninsula, and in eastern Africa, southwards to Tanzania.

It is one of the plants, which in Ancient Greece and Rome were utilized for production of a resin, known as silphium, silphion, or laser, used in perfume and medicine, and also as an aphrodisiac and a contraceptive. It was an essential trade item from the ancient North African city of Cyrene, and it was so important to the Cyrenian economy that most of their coins bore a picture of the plant. The gum is still produced in Morocco today.

It comes in two types, one poisonous to livestock, and one non-toxic. Young stems and inflorescences of the non-toxic type are edible. They were a food item in Ancient Rome, and are still eaten in Morocco.

The specific name is Latin, meaning ‘common’.

Pictures, depicting flowering specimens of this impressive plant, are shown on the page In praise of the colour yellow.

 

 

Giant fennel, growing along an abandoned railway, San Cataldo, Sicily. (Photo copyright © by Kaj Halberg)

 

 

 

Apocynaceae Dogbane family

 

Calotropis
A small genus with 3 species, distributed in the major part of Africa, and from the Near East and Arabia eastwards to southern China, southwards to Yemen, Sri Lanka, and Indonesia.

The fruit of one member, the Sodom apple (Calotropis procera), was first described by Roman historian Titus Flavius Josephus, born Yosef ben Matityahu (c. 37-100 A.D.), who found it growing near the city of Sodom: “which fruits have a colour as if they were fit to be eaten, but if you pluck them with your hands, they dissolve into smoke and ashes.” (W. Whiston, 1737. The War of the Jews, Book IV, Chapter 8, Sec. 4). Titus is referring to the fact that the ripe fruit easily dissolves, releasing hundreds of downy seeds, which are scattered to the four winds.

The generic name is derived from Ancient Greek kalos (‘beautiful’) and tropis (‘keel of a ship’), referring to the beauty of the flowers, and to the 5 keel-shaped ridges in the corona.

 

Calotropis gigantea Giant milkweed
This striking shrub may grow to 4 m tall, stem erect, branched, with a milky latex, leaves broadly elliptic or oblong, to 20 cm long and 12 cm broad, often stem-clasping, diminishing in size up the stem. The inflorescences are clusters from the leaf axils, flowers waxy, purplish-white with a lilac centre, to 4.5 cm across, petals 5, spreading star-like.

It is distributed from the Indian Subcontinent eastwards to southern China, and thence southwards to Indonesia, growing in disturbed habitats, including roadsides, fallow fields, and beaches.

The plant is widely used in traditional and ayurvedic medicine, described on the page Plants: Plants in folklore and poetry. The flowers are sometimes used as decoration. It is told that Hawaiian Queen Liliuokalani (1838-1917) wore them as a symbol of royalty, strung into leis (garlands).

 

 

Giant milkweed, growing along a railroad track in the city of Janakpur, southern Nepal. (Photos copyright © by Kaj Halberg)

 

 

 

Asteraceae Daisy family

 

Artemisia Mugwort, wormwood, sagebrush
These plants constitute a huge, worldwide genus, comprising between 200 and 400 species.

The genus is named for Artemis, the Greek goddess of wilderness and wild animals, hunting, childbirth and virginity, protector of young girls, and also bringer and reliever of disease in women. In his Herbarium, Roman philosopher and scholar Lucius Apuleius (c. 124-170 A.D.) writes: “Of these worts that we name Artemisia, it is said that Diana found them and delivered their powers and leechdom to Chiron the Centaur, who first from these worts set forth a leechdom, and he named these worts from the Greek name of Diana, Artemis.”

The role of Artemisia species in folklore and traditional medicine is described on the page Plants: Plants in folklore and poetry.

 

Artemisia vulgaris Common mugwort
This plant is native to the major part of Temperate Europe and Asia, and also to North Africa and Alaska. In other areas of North America, it has become naturalized. Originally, it was restricted to dry grasslands and sandy beaches, but when farming was introduced, it readily spread to fields, and today it is regarded as a noxious weed.

In China, this species is sometimes used as a substitute for A. argyii to make moxa, which is much utilized as a healer in traditional Chinese medicine.

The specific name is Latin, meaning ‘common’.

 

 

Common mugwort, Skanderborg Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cirsium Thistle
A huge genus of spiny herbs with about 400 species, distributed in temperate and subarctic regions of Eurasia, northern and eastern Africa, and Central and North America, southwards to Columbia, Zambia, the Malay Peninsula, and the Philippines.

These plants were already regarded as being different from the present-day members of the genus Carduus by Swiss botanist Gaspard Bauhin (1560-1624). He may have noticed that their pappus (seed hairs) form a plume, as opposed to members of Carduus, which have simple, unbranched seed hairs.

The generic name is derived from Ancient Greek kirsion, according to Greek physician, pharmacologist, and botanist Pedanius Dioscorides (died 90 A.D.) the name of a kind of thistle, derived from kirsos (‘swollen vein’), alluding to its usage against swollen veins.

A number of species in this genus are described on the page Plants: Thistles.

 

Cirsium arvense Creeping thistle, Canada thistle
This proliferous spiny herb, to 1.6 m tall, is found in most temperate areas of Asia and Europe, and has also been accidentally spread elsewhere around the globe.

It may be separated from other thistles by the rather small purplish, violet, or pink (rarely white) flowerheads, to 2 cm across. Flowering takes place between March and August.

In Nepal, a paste of the root is used for indigestion.

This species is a most troublesome weed in most cooler parts of the world. As far back as 1863, Danish research showed that a large creeping thistle can produce about 80 flowerheads in one season, and as flowerheads on average contain 120 flowers, one single plant is able to produce about 10,000 seeds per year! Its invasiveness is described on the page Nature: Invasive species.

The specific name is Latin, meaning ‘growing in fields’.

 

 

Creeping thiste and dyer’s chamomile (Cota tinctoria, see below), Copenhagen Central Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Creeping thiste in the fruiting stage, Aarhus Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cirsium vulgare Spear thistle
This extremely spiny plant, also known as bull thistle, is widely distributed in the major part of Europe, North Africa, and western Asia, eastwards to the Yenisei River. It has also been accidentally introduced to many other places, including North America and Australia, where it is often regarded as an invasive weed.

Spear thistle is biennial. During the first year, it produces a leaf rosette, which overwinters and grows into a stem, up to 1.5 m tall, the following summer, displaying an abundance of beautiful red flowerheads. Despite its formidable armour of spines, this plant has been elected as the national flower of Scotland, hence another common name, Scottish thistle. Other lovers of the spear thistle include honey bees, bumble bees, and butterflies, which feed on the nectar, and various finches, including goldfinch (Carduelis carduelis), linnet (Linaria cannabina), and greenfinch (Chloris chloris), which greatly appreciate the seeds.

The specific name is Latin, meaning ‘common’.

Several pictures, depicting this species, are shown on the page Plants: Flora in Turkey.

 

 

Spear thistle, Ringsted Railway Station, Zealand, Denmark. Horsetail fleabane (Erigeron canadensis, see below) is seen in the background. (Photo copyright © by Kaj Halberg)

 

 

 

Cota
Members of this genus, comprising about 36 species, were previously placed in the genus Anthemis (dog-fennels). They are native from Europe eastwards to eastern Siberia, southwards to the Mediterranean, Iran, and Pakistan.

According to one source, the generic name is derived from Ancient Greek cotyle (‘cavity’, ‘bowl’, or ‘cup’), alluding to the sessile leaves of some members of the genus forming a cavity at the base.

 

Cota tinctoria Dyer’s chamomile
This species, previously called Anthemis tinctoria, resembles the well-known corn daisy (Glebionis segetum), but may immediately be identified by its pinnately cut leaves.

Its native area is from England and Norway eastwards to eastern Siberia, southwards to the Mediterranean, Jordan, Iran, and Central Asia, growing on meagre soils, such as sandy fields and waste plots.

The specific name is Latin, meaning ‘used in dyeing’, derived from tingo (‘soak in dye’).

 

 

Dyer’s chamomile, Copenhagen Central Station, Denmark. In the upper picture, long-headed poppy (Papaver dubium), creeping thistle (Cirsium arvense), common mugwort (Artemisia vulgaris), and the reddish form of Herb Robert (Geranium robertianum) are also present, all described on this page. (Photo copyright © by Kaj Halberg)

 

 

 

Crepis Hawk’s-beard
This large genus, comprising about 200 species, is distributed in the Northern Hemisphere and Africa, with the core area around the Mediterranean.

The generic name is derived from Ancient Greek krepis (‘slipper’ or ‘sandal’), according to some authorities referring to the shape of the fruit.

 

Crepis capillaris Smooth hawk’s-beard
A widespread plant, native to the major part of Europe, eastwards to the Ural Mountains and the Caucasus, but widely introduced elsewhere, including North America, the northern Andes, South Africa, and Australia. It grows to 60 cm tall, with numerous pale yellow flowerheads to 1.5 cm across.

The specific name is derived from the Latin capillus (‘hair’ or ‘thread’), perhaps referring to the slender stem.

 

 

Smooth hawk’s-beard, Fredericia Railway Station, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Erigeron Fleabane
A huge genus with around 400 species, widespread in Asia, Europe, and North America, and also a few species in Africa and Australia.

These plants are very similar to some members of the well-known genus Aster, but ray florets are usually in 2 or several rows, and very narrow, as opposed to one row of broader florets in Aster.

The generic name is derived from Ancient Greek eri (‘early in the morning’) and geron (‘old man’), alluding to some species, which are covered with down when young. Some members of the genus contain an oil with a turpentine-like smell, which, supposedly, should deter fleas, hence the common name.

 

Erigeron canadensis Horsetail fleabane
This plant is also known as Canadian fleabane, and some authorities regard it as belonging to the genus Conyza. It is native to North America and parts of Central America, but has been accidentally introduced to large parts of the world. In many places it has become a serious pest, especially in Europe and Australia, but also in its native North America.

It prefers to grow in undisturbed areas and is particularly troublesome in newly established plantations, where it is able to resist herbicides, growing to 3 m tall, thus depriving planted species of nutrients and sunlight.

It contains an oil with a turpentine-like smell, which, supposedly, should deter fleas, hence its common name. Another popular name is bloodstanch, given by herbalists, who claim that an extract from leaves and flowers arrests haemorrhages from the lungs and alimentary tract.

 

 

Horsetail fleabane, Ringsted Railway Station, Zealand (top), and Skanderborg Railway Station, Jutland, both Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Hypochaeris Cat’s-ear
This genus is distributed in temperate areas of Europe, Asia, Africa, and South America. The number of species is disputed, varying from about 64 to 100, depending on authority.

The generic name, previously written Hypochoeris, is probably derived from Ancient Greek hypo (‘under’) and choeris (‘young pig’), alluding to the fact that pigs enjoy eating the root of common cat’s-ear (below). The English name refers to the leaves of this species, which are covered in rough hairs.

 

Hypochaeris radicata Common cat’s-ear
Common cat’s-ear is native to the major part of Europe and northern Africa, but has been introduced to many other parts of the world, including the Americas, Japan, Australia, and New Zealand. In some areas, it is considered an invasive weed. It grows in grasslands, along roads and banks, in old dunes, and in waste places.

It is a perennial herb, the forked stem sometimes growing as tall as 80 cm. The basal leaves, which form a rosette, are obovate in outline, to 20 cm long and 4 cm wide, margin wavy or lobed, sometimes pinnately divided, hairy, stem leaves absent or few, similar to the basal leaves. The terminal flowerheads are bright yellow, to 1.5 cm across.

A close relative, large cat’s-ear (H. uniflora) is described on the page Plants: Flora in the Alps and the Pyrenees.

 

 

Common cat’s-ear, growing together with the red form of Herb Robert (Geranium robertianum) and common goat’s-beard (Tragopogon pratensis) (both in front), and false oat-grass (Arrhenatherum elatius), Copenhagen Central Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Common cat’s-ear, Nyborg Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Jacobaea Ragwort, groundsel
A genus with about 66 species, found in all of Europe and most parts of Asia, in North Africa, and in northern parts of North America. They were formerly placed in the genus Senecio (below).

The generic name refers to Jacobus, in English called James the Great, one of the twelve apostles of Jesus. He is the patron saint of Spain and, according to tradition, his remains are housed in the city of Santiago de Compostela, Galicia, known as the culmination of the Camino de Santiago pilgrimage route.

The name ragwort was given in allusion to the ragged foliage of many of these plants.

Several members are described on the page Plants: Groundsels, ragworts, and allies.

 

Jacobaea vulgaris Tansy ragwort, common ragwort
This biennial or perennial, previously known as Senecio jacobaea, is native to the entire Europe, southwards to North Africa and Turkey, and thence eastwards to the Ural Mountains and Central Asia. It has also been introduced accidentally to many other places and is often considered a troublesome weed, mainly due to its toxicity to cattle and horses. It usually grows in dry grasslands and other open places.

The stem is erect, green or reddish, usually smooth, much branched, sometimes reaching a height of 2 m, but usually lower. The leaves, to 20 cm long and 6 cm wide, are pinnately lobed, the lobes toothed along the margin. The very numerous flowerheads, to 2.5 cm across, have 10-15 yellow, strap-shaped ray florets and many orange disc florets. The flowers are much visited by bees, hover flies, and butterflies.

The specific name is Latin, meaning ‘common’.

The name tansy ragwort refers to the foliage, which somewhat resembles that of tansy (Tanacetum vulgare, see below). Popular names also include stinking willie and mare’s fart, alluding to the unpleasant smell of the leaves.

 

 

Common ragwort, Copenhagen Central Station, Denmark. The other plants are common mugwort (Artemisia vulgaris) and false oat-grass (Arrhenatherum elatius). (Photo copyright © by Kaj Halberg)

 

 

 

Lactuca Lettuce
The number of species in this genus is unsettled. Many former members have been moved to other genera, and the genus is still being revised. Kew Gardens accepts a total of 117 species, other sources mention 50-70 species. Members are mainly distributed in temperate areas of Asia, Europe, and North America.

The generic name was the classical Latin name of the garden lettuce (L. sativa), derived from lactis (‘milk’), alluding to the milky sap of this plant. The English name is a corruption of the Latin word.

 

Lactuca muralis Wall lettuce
This species, by some authorities called Mycelis muralis, is native to most of Europe, north-western Africa, and West Asia, eastwards to the Caucasus, where it may be found up to an altitude of 2,300 m. It has also become naturalized in North America and New Zealand.

The main habitat of this plant is woodlands, but it may also be found in open areas, including forest clearings, city walls, stone fences, and along railroads. The specific name is derived from the Latin murus (‘wall’).

 

 

Wall lettuce, growing between railroad tracks, Horsens, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Lactuca serriola Prickly lettuce
This plant is named for the row of spines along the mid-vein on the underside of the leaves, and the leaf margins also have fine spines. The leaves are very variable, from entire to deeply divided. It is a tall plant, under favourable conditions growing to 2 m tall, with a slightly fetid smell.

It is native to Europe, North Africa, and temperate areas of Asia, and has also become naturalized elsewhere, growing along beaches, roads, and railroads, and as a field weed.

One folk name of prickly lettuce is compass plant. The leaves of its main stem are aligned north-south, offering the least surface area to the midday sun, but the maximum area to the weaker light early and late in the day.

Incidentally, this plant was named twice by Swedish naturalist Carl Linnaeus (1707-1778), in 1756 as L. serriola, and in 1763 as L. scariola, derived from the Latin escarius (‘edible’). The first name is probably a misspelling, which Linnaeus then corrected in 1763. However, according to the nomenclature rules, the first published name has priority.

The name scariola was used by Englishmen at least as far back as the 1400s. According to the Oxford Dictionary, one source said: “Wylde letus hat feldman clepyn Skariole.” (“Wild lettuce have field-men (farmers) called Skariole”).

 

 

Prickly lettuce, growing at Ringsted Railway Station, Zealand, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Lapsana communis Nipplewort
This plant, growing to about 1.2 m tall, is native from Europe eastwards to central Siberia, southwards to northern Africa, Iran, and Pakistan, but has become naturalized elsewhere, including North America and Australia. According to Kew Gardens, it is the only member of the genus.

It grows in forests, hedges, along roads and railroads, and in fallow fields. The young leaves are edible, cooked like spinach or eaten raw in salads.

The common name stems from the 17th century, when the word nipple was regarded as being equivalent to the word papillaris, derived from the Latin papilla (‘nipple’). In Germany, followers of the Doctrine of Signatures maintained that the plant could be used to treat cracked nipples, as its flowerheads, when young, resemble nipples.

The specific name is Latin, meaning ‘common’.

 

 

Nipplewort, growing along abandoned railway tracks, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Matricaria Chamomile, mayweed
A small genus with 6 species, widely distributed in the Northern Hemisphere.

The generic name is derived from the Latin matricis (’mother’s life’, i.e. the womb). Formerly another plant, feverfew (Tanacetum parthenium), was regarded as a species of chamomile, named Matricaria parthenium by Swedish naturalist Carl Linnaeus (1707-1778). In Ancient Rome, feverfew was used for uterus problems.

 

Matricaria discoidea Disc mayweed
Disc mayweed is native to North America and north-eastern Asia, southwards to Hokkaido, Japan. Today, however, it has been accidentally introduced to most other areas of the world, where it is a common weed, growing in open areas. Other names of the plant include wild chamomile and pineapple weed, due to its chamomile- or pineapple-like smell, when crushed.

The specific name is derived from the Latin discoides (‘disc-shaped’), referring to the flowerheads.

 

 

Disc mayweed, Nyborg Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Senecio Ragweed, groundsel
Formerly, this genus was huge, but many species have been transferred to other genera. However, it is still very large, comprising more than 1,400 species, found almost worldwide. The majority are erect herbs, with a few climbing or scrambling species.

The generic name is derived from the Latin senex (‘old man’), alluding to the white seed hairs of the genus.

The name ragwort was given in allusion to the ragged foliage of many of these plants.

Several members are described on the page Plants: Groundsels, ragworts, and allies.

 

Senecio vernalis Eastern groundsel
This annual blooms early in the year. It may grow to 60 cm tall, stem simple or branched above, often covered in woolly hairs, but may also be smooth. The leaves are pinnately divided, often woolly-haired, to 10 cm long and 4 cm broad, lobes toothed. Inflorescences are terminal, usually with few flowerheads, each to 2.5 cm across. Ray florets, usually around 8, are bright yellow, to 9 mm long, disc florets orange-yellow.

Its habitats include roadsides, fields, waste places, inner parts of beaches, and other open, disturbed areas.

Originally, it was a native of south-eastern Europe and western Asia, but was accidentally introduced to other parts of Europe in the 1700s, presumably with imported crops. Today, it is extremely common, especially on poor soils, and is regarded as a severe pest in grass, clover, and alfalfa fields.

The specific name is Latin, meaning ‘of the spring’, derived from ver (‘spring’).

 

 

Eastern groundsel, Copenhagen Railway Station, Denmark. A number of other plants are also seen, including corn salad (Valeriana locusta), dandelion (Taraxacum officinale), colt’s foot (Tussilago farfara), soft brome (Bromus hordeaceus), and snowdrop anemone (Anemone sylvestris), the latter an escape from a garden, as it does not grow in the wild in Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Senecio viscosus Sticky ragwort, sticky groundsel
As its name implies, this plant, also called stinking groundsel, is very sticky. The following quotation gives a vivid impression of its stickiness: Sticky groundsel has characteristic glandular hairs, which secrete a substance that is as sticky as fly paper, and by the end of summer it is quite a mess with all the dust, sand, small insects, hairs, feathers, downy seeds, its own cypselas [seeds], candy wrappers, and who knows what else that have stuck to it.” (Source: luontoportti.com/suomi/en/kukkakasvit/sticky-groundsel)

It is very similar to the common woodland ragwort (S. sylvaticus), which, however, does not have sticky glandular hairs. Sticky ragwort also has longer ray florets, which are initially spreading, later recurved.

Originally, sticky ragwort was a native of southern and central Europe and western Asia. Since the 1800s, it has spread considerably and has become naturalized in northern Europe, Canada, United States, and elsewhere.

The specific name is Latin, meaning ‘sticky’, derived from viscum, the classical word for a birdlime, made from berries of the common mistletoe (Viscum album).

 

 

Sticky groundsel, Aarhus Railway Station, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Sonchus Sow-thistle
This genus, comprising about 100 species, is widely distributed in the entire Old World, and has also become widely naturalized in the New World.

The generic name is a Latinized form of sonkhos, the ancient Greek word for sow-thistles. The common name refers to the fact that pigs like to eat these plants, and to the leaves, which resemble young thistle leaves.

A number of species are described on the page Plants: Urban plant life.

 

Sonchus asper Prickly sow-thistle
A stout plant, which may grow to 1.2 m tall, but is often much lower, often with a reddish stem. The leaves are extremely variable, obovate, spatulate, or elliptic, to 13 cm long and 5 cm wide, entire or irregularly divided, base eared and strongly recurved, clasping the stem, margin heavily spiny. Flowerheads are relatively few, to 2.5 cm across, in an open terminal cluster, ray florets numerous, bright yellow, disc florets absent.

This species presumably originates from the Mediterranean region, but has become naturalized in most parts of the world. In Nepal, a paste of the plant is applied to wounds and boils. It is also collected for fodder, and tender parts are cooked as a vegetable.

The specific name is Latin, meaning ‘rough’ or ‘coarse’, naturally alluding to the leaves.

 

 

Prickly sow-thistle, Silkeborg Railway Station, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Sonchus oleraceus Common sow-thistle
It is told from the previous species by its more slender stem, occasionally to 1.5 m tall, and much less spiny leaves, which may grow to 20 cm long and 9 cm wide. The base of the leaf is not eared, but has two straight, lanceolate lobes, which clasp the stem. The flowerheads are smaller, conical, to 2 cm across, ray florets pale yellow, disc florets absent. The flowering period is long, from May to October.

It is probably native to Europe and western Asia, but has been spread to most other areas of the world, growing in waste places, fallow fields, along roads, and in cities. It is regarded as an invasive plant in many countries, including Australia, where it is a serious problem in crops.

The specific name is derived from the Latin oleris, meaning ‘edible’. Young leaves can be eaten as salad or cooked like spinach. The plant is also used medicinally.

 

 

Common sow-thistle, growing along a railroad track in the city of Aarhus, Denmark. The red plants in the background are the red form of Herb Robert (Geranium robertianum), and the green plant is tufted vetch (Vicia cracca). (Photo copyright © by Kaj Halberg)

 

 

 

Tanacetum Tansy
This genus, comprising about 100 species, is distributed in Europe, North Africa, the Middle East, and Central Asia, and northern North America. About 36 species, which were formerly placed in this genus, have been moved to the genus Ajania.

Disc florets are yellow, whereas ray florets are missing in most species. When present, they are mostly white.

The common name was adapted from Old French tanesie, which came from the Latin tanacetum, which was again adapted from Ancient Greek athanatos (‘immortal’), according to some old times herbalists referring to the conserving properties of common tansy (below), which is toxic to various pests, blocking their attack.

 

Tanacetum vulgare Common tansy
This plant is widely distributed in Eurasia, from England and Spain eastwards to the Pacific, and from subarctic areas southwards to the Mediterranean, Turkey, Tibet, and Japan. In former times, it was much utilized in folk medicine, but today it is mainly used for dyeing.

The specific name is Latin, meaning ‘common’.

Other pictures, depicting this species, are shown on the page In praise of the colour yellow.

 

 

Common tansy, Aarhus Railway Station, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Taraxacum Dandelion
Apomictic (asexual) reproduction is very common in dandelions, resulting in an abundance of species and micro-species, more than 2,500 worldwide, mainly in the Arctic and temperate zones of the Northern Hemisphere, a few in temperate regions of the Southern Hemisphere. They grow in a variety of open areas, including waste places, fallow fields, along roads, and in cities.

These plants are characterized by stem and leaves having a milky sap. The long and narrow leaves are in a rosette, 10-20 cm across, blade usually lobed or serrated, but sometimes almost entire, wavy. The single flowerhead, 2-6 cm across, is borne on a leafless, hollow stem, 10-20 cm tall, sometimes more. Florets yellow, very numerous, densely crowded. Flowering takes place most of the year.

Dandelion was first mentioned as a medicinal herb by Arabian physicians in the 10th or 11th Century, who called it a sort of wild endive (Cichorium), under the name of tharakhchakon, which was corrupted to Taraxacum. In French, these plants were named dent de lion (‘lion’s tooth’), alluding to the often strongly serrated leaves. This name was adopted by the English, corrupted to dandelion.

 

Taraxacum officinale Common dandelion
This plant is ubiquitous everywhere in Europe, also along railroad tracks, and it has been spread to many other areas of the world. It is described in depth on the page Plants: Plants in folklore and poetry.

 

 

Common dandelion, Alvesta Railway Station, southern Sweden. Also seen are young plants of common sow-thistle (Sonchus oleraceus) (left of the dandelion) and prickly lettuce (Lactuca serriola) (upper left corner). (Photo copyright © by Kaj Halberg)

 

 

 

Tussilago farfara Colt’s-foot
The only species in the genus, distributed all over Europe, North Africa, Turkey, the Caucasus, northern Iran, Central Asia, the Himalaya, and China. It is widely naturalized in North America.

Flowering takes place early in spring, from February to April, long before the leaves develop. The flowering stems, to 10 cm tall, elongated in fruit, are covered in whitish down, with scaly, purplish bracts. Flowerheads are solitary, terminal, to 3 cm across, nodding before and after flowering. The involucre is bell-shaped, covered in white down, to 1.8 cm long at fruiting. The bright yellow ray florets are very numerous, narrow, to 1 cm long, disc florets orange-yellow.

The leaves, all basal, appear after the flowers have withered. They are long-stalked, green above, white-woolly below, very broadly heart-shaped, almost circular in outline, to 14 cm across, margin wavy, with many or few teeth.

Leaves and young flowerheads are used medicinally for cough. In China, the plant is also cultivated as a honey plant.

The generic name, derived from the Latin tussis (’to cough’), refers to the usage of the leaves. The specific name is derived from farfarum, the classical Latin name of the plant. The common name was given in allusion to the shape of the leaves. An old name for the plant was Filius ante patrem (‘the son before the father’), referring to the fact that the flowers appear long before the leaves.

 

 

Leaves of colt’s-foot, together with false oat-grass (Arrhenatherum elatius), Aarhus Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Balsaminaceae

 

Impatiens Balsam, touch-me-not, jewelweed
These plants are characterized by their fleshy, almost translucent stems, and the unique structure of their attractive flowers, having 3 or 5 sepals, of which the lower one is greatly enlarged, funnel- or pouch-shaped, with a spur, whereas the other 2 or 4 are small and greenish. There are 5 petals, of which the upper one, the standard, is often helmet-like, whereas the 4 lateral ones are fused in pairs, the upper pair forming the wings, the lower pair the lip.

The generic name, as well as the popular name touch-me-not, were given in allusion to the way these plants spread their seeds. As the fruit reaches maturity, a tension builds up inside the pod, causing it to ‘explode’ when touched, hereby spreading the seeds a considerable distance.

The common name balsam is in fact a misnomer. It was adopted from the garden balsam (Impatiens balsamina), which was brought to Europe by the Portuguese in the 16th Century. Originally, balsamine was a Greek name of an unidentified plant with an aromatic oil, derived from balsamon (‘balsam’), which in the Hebrew means ‘spice’ or ‘perfume’.

 

Impatiens parviflora Small-flowered balsam
The original range of this plant is probably mountains of Central Asia, but it has become widely naturalized elsewhere. It is regarded as an unwanted weed in many countries, often dispelling native plants in damp and shady forests.

When cooked, the leaves are edible, and the seeds can be consumed raw or cooked. Medicinally, it has been used for treatment of warts, ringworm, and nettle stings, and also to relieve an itchy scalp.

The specific name is Latin, meaning ‘with small flowers’.

 

 

Small-flowered balsam, Copenhagen Railway Station, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Boraginaceae Borage family

 

Myosotis Forget-me-not
This large genus with about 150 species is found in Asia, Europe, Africa, Australia, and North America.

The generic name is derived from Ancient Greek myos (’mouse’) and otis, which is the nominative plural of ous (’ear’), thus ‘mouse-ears’, alluding to the shape and hairiness of the leaves.

The origin of the name forget-me-not, and the role of these plants in folklore, is described on the page Plants: Plants in folklore and poetry.

 

Myosotis arvensis Field forget-me-not
A most annoying weed in fields, as its seeds may retain their ability to sprout for about 30 years.

It is native to temperate areas of Eurasia, from Iceland eastwards to eastern Siberia, southwards to northern Africa, Turkey, and Kazakhstan. It has become naturalized in many other places, including New Zealand and North America, where it sometimes becomes invasive.

The specific name is Latin, meaning ‘growing in fields’.

 

 

Field forget-me-not, growing at Nyborg Railway Station, Funen, Denmark. Prickly-headed poppy (Papaver argemone) and prickly lettuce (Lactuca serriola) are also present. (Photo copyright © by Kaj Halberg)

 

 

 

Brassicaceae Cabbage family

 

Alliaria
This small genus contains only 2 species, A. petiolata (below), and A. taurica, which is restricted to the Caucasus.

The generic name is derived from Allium, the genus name of onions and garlic, referring to the garlic-like smell of crushed leaves of these plants.

 

Alliaria petiolata Garlic mustard
This plant is native from northern and western Europe eastwards to western Siberia and Xinjiang, southwards to north-western Africa, Iran, and northern Pakistan.

Around 1860, it was introduced to the United States as a spice herb, and since then it has spread to most states, and to Canada as well. It has invaded numerous forests, where it is able to dominate the understorey, expelling native plants. During the last 50 years, it has also been spreading considerably in its native area, presumably benefitted by the increased nitrogen deposition.

It is biennial, in the first year forming a leaf rosette at ground level. It remains green through the winter, developing a stem the following spring, to 1 m tall, occasionally to 1.3 m. Leaves are stalked, broadly triangular or heart-shaped, blade to 8 cm long and across, margin toothed. Inflorescences are small, long-stalked clusters of white flowers from the leaf axils, petals to 8 mm long and 3 mm wide.

The specific name is Latin, means ‘having petioles’, i.e. leaf-stalks.

 

 

Leaves of garlic mustard, growing at Nyborg Railway Station, Funen, Denmark. To the right cleavers (Galium aparine, see below). (Photo copyright © by Kaj Halberg)

 

 

 

Arabidopsis Rock-cress
The native area of this genus, comprising about 12 species, is temperate areas of the Northern Hemisphere, and also mountains in tropical areas of eastern Africa.

The generic name is composed of the genus name Arabis, also in Brassicaceae, and Ancient Greek opsis (‘appearance’ or ‘resemblance’), thus ‘resembling Arabis‘.

 

Arabidopsis arenosa Sand rock-cress
This species is a native of central Europe, from France eastwards to Latvia and Ukraine, and from south-eastern Denmark south to northern Italy and Macedonia, but has been introduced accidentally elsewhere.

Stem erect, to 30 cm long, often branched from the base, lower part hairy. Basal leaves form a rosette, blade obovate or oblanceolate, pinnately divided, with 3-11 lobes on each side, to 5 cm long and 1 cm wide, stem leaves smaller, usually entire. Flowers to 1 cm across, petals white or lavender.

The specific name is derived from the Latin harena (‘sand’), earlier hasena, possibly from Etruscan, and the suffix osus (‘full of’). Thus an approximate meaning is ‘abundant in sand’.

 

 

In Sweden, sand rock-cress is very common, mainly spreading along railways. This picture is from Alvesta Railway Station, in the southern part of the country. (Photo copyright © by Kaj Halberg)

 

 

 

Barbarea Wintercress
Members of this genus, containing about 30 species, are found in temperate areas of the Northern Hemisphere, and also on mountains in tropical areas of eastern Africa.

The generic name is adapted from a medieval name of the plant, Herba Sanctae Barbarae (’Saint Barbara’s wort’). Holy Barbara was regarded as the patron saint of artillery men and miners, and as wintercress was considered a good healer of wounds, it was named after her.

According to legend, Barbara was the daughter of a rich pagan, living in the 3rd Century. Her father kept her locked up in a tower in order to protect her from the influence of the outside world. However, she secretly became a Christian and rejected to marry the man her father presented to her.

Before going on a journey, her father ordered a private bath-house to be constructed for her near her dwelling, but during his absence, Barbara had three windows put in it, as a symbol of the Holy Trinity, instead of the two originally intended. When her father returned, she informed him that she had become a Christian. He dragged her before the prefect of the province, but, despite being cruelly tortured, she held true to her Christian faith.

At night, her prison was bathed in light, and every morning her wounds were healed. Torches that were held near her to burn her were extinguished. Finally, she was beheaded, and her father himself carried out the death sentence. As punishment, he was struck by lightning and his body consumed by flame.

Barbara was buried by a Christian, and her tomb became the site of miracles. In his book Reisebeschreibung nach Arabien und andern umliegenden Ländern (Gloyer und Oldshausen, Hamburg, 1837, in English Travels Through Arabia and Other Countries in the East, Performed by M. Niebuhr, Gale Ecco, 2018), German cartographer and mathematician Carsten Niebuhr relates that he was shown her grave in the church of the village Karmelis, near Mosul, northern Iraq, in 1766. Some nearby ruins were said to have been her father’s palace.

 

Barbarea vulgaris Common wintercress
Today, this yellow-flowered plant, also known as yellow rocketcress, winter rocket, or wound rocket, is very common in northern Europe, especially in disturbed places, including abandoned plots, and along roads and railways. This was not always the case, as it was introduced during the 1700s from its native areas in southern Europe, North Africa, and the Near East. It has also been introduced to many other places, including Iceland, North America, and Australia.

The specific name is Latin, meaning ‘common’.

 

 

Common wintercress, Alvesta Railway Station, southern Sweden. A small dandelion (Taraxacum officinale) is seen to the right. (Photo copyright © by Kaj Halberg)

 

 

 

Campanulaceae Bellflower family

 

Campanula Bellflower
This genus includes more than 500 species, found in temperate and subtropical areas of the Northern Hemisphere, with the highest diversity around the Mediterranean and in the Middle East. No less than 44 species occur in the Alps.

The generic name and the popular name, as well as the name of the entire family, were applied in allusion to the bell-shaped flowers of this genus, from the Latin campanula (’little bell’), the ‘bell’ consisting of five fused petals with free tips. This name was first used in 1542 by German physician and botanist Leonhart Fuchs (1501-1566).

 

Campanula rapunculoides Creeping bellflower
This plant grows to 80 cm tall, occasionally to 1.2 m. The stem is simple, erect, slightly downy. The basal leaves are narrowly triangular, with a heart-shaped or rounded base, toothed, to 12 cm long and 2 cm wide, upper stem leaves lanceolate, smaller. The inflorescence is a long, one-sided raceme with numerous short-stalked, drooping flowers, blue or violet-blue, rarely white, lobes slightly hairy, sepals narrow, back-curved, bracts small, pointed, leaf-like.

Its habitats include open forests, shrubberies, fields, gardens, and along roads, railways, and hedgerows, preferably in partial shade. It spreads by underground rhizomes, often forming dense stands. Even a small piece of rhizome can spout into a new plant, making it very hard to eradicate, once it has entered a garden.

It is found in all of Europe, except Iceland and the British Isles, southwards to the Mediterranean, Turkey, and Iran, eastwards to central Siberia and Central Asia. It has also been introduced to North America, where it has become an extremely invasive weed.

The specific name refers to the similarity of this plant to C. rapunculus, whose specific name is a diminutive of the Latin rapa (‘turnip’), thus ‘little turnip’, referring to the shape of the root. C. rapunculus is described on the page Plants: Plants in folklore and poetry.

Other bellflower species are presented on the pages Plants: Flora of the Alps and the Pyrenees, and In praise of the colour blue.

 

 

Creeping bellflower, growing at a fence around Alvesta Railway Station, southern Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Caprifoliaceae Honeysuckle family

 

Valeriana Valerians
A huge genus with about 425 species, found worldwide, except in polar regions and some areas in Africa, tropical Asia, and Australia.

Previously, these plants were placed in the family Valerianaceae, which has now been reduced to a subfamily, Valerianoideae, of the honeysuckle family.

The generic name is derived from the Latin valere (‘to be strong, healthy’), alluding to the medicinal properties of many valerian species.

 

Valeriana locusta Corn salad, lamb’s lettuce
This small annual, which was previously placed in the genus Valerianella, is native to Europe, western Asia, and northern Africa, and it is also widely cultivated as a vegetable. In North America it has escaped cultivation and become naturalized in many places. Its habitats include waste areas, inner sandy beaches, and along railways, and it is a common weed in farmland.

It forms a basic leaf rosette, blade spatulate, to 5 cm long, stem leaves smaller, to 2 cm long, stems several, branching out from the rosette, to 35 cm long. Inflorescences are small, branched clusters from the leaf axils, corolla bluish-white, with 5 fused petals, to 2 mm long and wide.

In a botanical context, the specific name means a ‘spikelet in a flower cluster’. What it refers to in this case is not clear.

 

 

Corn salad, growing at Nyborg Railway Station, Funen, Denmark. In the upper picture, prickly-headed poppy (Papaver argemone) and hairy vetch (Vicia hirsuta) are also present. (Photos copyright © by Kaj Halberg)

 

 

 

Valeriana rubra Red valerian
This plant, also known as Centranthus ruber, has a branched stem, often woody at the base, to 80 cm tall, erect or ascending, leaves sessile, fleshy, bluish or greyish-green, ovate or broadly lanceolate, long-pointed, to 10 cm long and 3 cm wide, margin entire or toothed. The inflorescence is a dense terminal cluster of fragrant, spurred flowers, corolla bright red, pink, or white, tube to 9 mm long, spur to 7 mm.

It grows in a variety of habitats, including dry, sandy or rocky places, and on walls, from sea level up to altitudes around 1,000 m. It is native from western Europe and Morocco eastwards to Turkey, but has been introduced to many other places as a garden plant. It has escaped and become naturalized in many countries, including the British Isles, Australia, South Africa, and the United States, and is regarded as invasive in the latter two countries.

The specific name is Latin, meaning ‘red’, alluding to the flower colour.

 

 

Red valerian, growing along an abandoned railroad track, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Caryophyllaceae Carnation family

 

Arenaria Sandwort
A huge genus with more than 300 species, found in northern temperate, subarctic, and arctic regions. No less than 102 species occur in China, mainly in arid areas.

The capsule of these plants has 6 teeth.

The generic name is derived from the Latin harena (‘sand’), earlier hasena, possibly from Etruscan. The name refers to the fact that many members of the genus often grow in sandy areas, which is also reflected in the English name.

 

Arenaria serpyllifolia Thyme-leaved sandwort
The stems of this many-branched annual are ascending or creeping, to 30 cm long, densely hairy. The leaves are sessile, ovate, to 1.2 cm long and 7 mm broad, smooth or sparsely hairy, tip pointed, margin hairy. Inflorescences are many-flowered, spreading clusters, sepals 5, lanceolate, to 4 mm long, pointed, petals 5, white, obovate, shorter than sepals.

It is distributed in a vast area, from all of Europe (except Iceland) eastwards to the Ural Mountains, and from northern and eastern Africa across Arabia, Central Asia, and the Himalaya to Japan and Taiwan. It has also become widely naturalized in North and South America, Australia, and New Zealand.

It grows in sandy and stony places, as a weed in fields and gardens, on walls, and along roads. In Central Asia, it has been recorded up to elevations around 4,000 m.

In Asia, it is used medicinally.

The specific name is derived from Ancient Greek herpyllos, the word for a species of thyme (Thymus), which in Old Latin was corrupted to serpyllum, and the Latin folium (‘leaf’), thus ‘with leaves like thyme’.

 

 

Thyme-leaved sandwort, Nyborg Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cerastium Mouse-ear chickweed
A large genus with about 100 species, almost cosmopolitan, mostly in temperate and arctic areas.

The calyx has 10 teeth in most species.

The generic name is derived from Ancient Greek keras (‘horn’), alluding to the horn-shaped capsules, which protrude from the calyx.

 

Cerastium fontanum Common mouse-ear chickweed
This plant, branched from the base, has ascending or erect stems, to 45 cm long, basal leaves ovate, lanceolate, or spatulate, to 1.3 cm long and 1 cm broad, hairy, stalked, stem leaves sessile or very short-stalked, ovate or oblong, to 3 cm long and 1 cm broad, tip pointed, margin densely hairy. Inflorescences are terminal, spreading, lax clusters with up to 40 flowers, sepals oblong, ovate, or lanceolate, to 6 mm long, margin often with a membrane, petals white, obovate or oblong, to 1 cm long.

It is distributed from Greenland, Iceland, and Scandinavia southwards to Morocco, eastwards to Finland and the Balkans. It has also been introduced to the Himalaya, Japan, and almost the entire North America.

It grows in open areas, including roadsides, trails, slopes, forest margins, and grasslands, and is also a weed in gardens and fields. In Central Asia, it has been recorded up to elevations around 4,300 m.

The specific name is derived from the Latin fons (‘spring’), indicating that the plant is fond of damp areas.

 

 

Fruiting common mouse-ear chickweed, Nyborg Railway Station, Denmark. An abundance of downy seeds from another plant are clinging to it. (Photo copyright © by Kaj Halberg)

 

 

Withered common mouse-ear chickweed, Copenhagen Railway Station, Denmark. The tiny green plant is procumbent pearlwort (Sagina procumbens, see below). (Photo copyright © by Kaj Halberg)

 

 

 

Sagina Pearlwort
This genus of small herbs, containing about 36 species, is found in subarctic, temperate, and subtropical regions on all continents, except Antarctica, and some members also occur in certain tropical mountains.

The generic name is Latin, meaning ‘feasting’, alluding to members of the genus Spergula (spurreys), which were once included in Sagina. Members of Spergula were utilized as vegetables.

The common name refers to the small, white, rounded flowers.

 

Sagina procumbens Procumbent pearlwort
This small plant is native from the entire Europe eastwards til central Siberia, southwards to northern Africa, Iran, and Xinjiang, but has been accidentally introduced to many other parts of the world. It is common in open places, including fields and along roads, and has also adapted to a life in cities.

The specific name is Latin, meaning ‘creeping’ or ‘lying flat on the ground’.

 

 

Procumbent pearlwort, Copenhagen Railway Station, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Spergularia Sand-spurrey
Another genus of small herbs, comprising about 66 species, found in temperate regions on all continents, except Antarctica, and some members also occur in certain tropical mountains.

The generic name is Latin, meaning ‘resembling Spergula’ (spurreys).

 

Spergularia rubra Red sand-spurrey
Red sand-spurrey is native to most of Europe, eastwards through Siberia to the Pacific coast, southwards to northern Africa, Ethiopia, Arabia, Xinjiang, and north-eastern China. It has also been introduced to many other countries.

It is common in open places like fields and along roads, and it may sometimes be encountered in cities.

The specific name is Latin, meaning ‘red’, presumably referring to the flower colour.

 

 

Red sand-spurrey, Skanderborg Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Convolvulaceae Morning-glory or bindweed family
Many members of this family are described on the page Plants: Morning-glories and bindweeds.

 

Calystegia False bindweed
This genus contains about 25 species of herbaceous twining vines, some of which may grow to 5 m long, or more. They are distributed in temperate and subtropical areas around the world, with almost half of the species being endemic to California. The fruit is a capsule.

The generic name is derived from Ancient Greek kalyx (‘cup’) and stegos (‘covering’), thus ‘a covering cup’, referring to the shape of the calyx.

Formerly, these plants were included in the genus Convolvulus (below), which they closely resemble, only differing in minute botanical characters.

 

Calystegia sepium Hedge bindweed
This species, divided into 9 subspecies, has a very wide distribution in the northern temperate zone, less so in the southern temperate zone.

The stem is twining in a counter-clockwise direction, reaching a length of up to 3 m. The leaves are alternate, dull green above, paler below, arrow-shaped, simple, to 10 cm long and 7 cm wide, flowers trumpet-shaped, to 7 cm long, white in most subspecies, although an American subspecies, americana, has a white and pink corolla, much like the cultivated hairy bindweed (C. pulchra). It is distributed across southern Canada and the eastern half of the United States, replaced by other subspecies further west.

Due to its vigorous growth, hedge bindweed is regarded as a pest in many places.

The specific name is Latin, meaning ‘growing in hedges’, alluding to this plant being common in hedges.

 

 

Hedge bindweed, growing along abandoned railway tracks, Nyborg, Denmark. Chicory (Cichorium intybus) is seen in the upper right corner. (Photo copyright © by Kaj Halberg)

 

 

 

Convolvulus Bindweed
Bindweeds, comprising about 190 species, are distributed across the major part of the planet, with the greatest diversity in the Middle East. Most species are twining herbs, although quite a few are woody shrubs or dwarf shrubs. The fruit is a capsule.

The generic name is derived from the Latin convolvere (‘to wind’ or ‘to wrap’ (around)), ultimately from con (‘with’) and volvo (‘to roll’).

 

Convolvulus arvensis Field bindweed
While most bindweeds twine around other plants, this species, growing to 2 m long, often grows along the ground.

It is native to temperate and subtropical areas of the Old World, from Ireland eastwards to eastern Siberia, and from Scandinavia and Siberia southwards to northern Africa, India, and northern Indochina. It has been accidentally introduced to many other parts of the world, where it easily becomes naturalized, and has become invasive in many places. This issue is described on the page Nature: Invasive species.

In America, where this plant is mainly known as Creeping Jenny, many of its popular names allude to its invasive nature. Edwin Spencer (see above) says: “Creeping Jenny is one of the meanest of weeds. That name aptly describes it. A whispering little hussy that creeps in and spoils everything. The weed needs no other name than this, but it has several others (…) hedge bells, corn-lily, withwind, bellbine, lap-love, sheep-bine, corn-bind, bear-bind, and green vine.”

The specific name is derived from the Latin arvus (‘cultivated’), thus ‘growing in cultivated fields’.

 

 

Field bindweed, Alvesta Railway Station, southern Sweden. The plant to the left is yellow dock (Rumex crispus). (Photo copyright © by Kaj Halberg)

 

 

Field bindweed, growing along abandoned railway tracks, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Dryopteridaceae Woodfern family
 
Dryopteris Woodferns
A huge genus with about 350 species, found on all continents except Antarctica.

The generic name is derived from Ancient Greek drys (‘oak’) and pteris (‘fern’), originally the classical Greek name of the black spleenwort (Asplenium adiantum-nigrum).

 

Dryopteris filix-mas Common male fern
This well-known fern is found throughout northern temperate areas in Eurasia and North America. Various forms are cultivated as ornamentals.

Greek scholar Theophrastos (c. 371-287 B.C.) and Greek physician, pharmacologist, and botanist Pedanius Dioscorides (died 90 A.D.), who was the author of De Materia Medica, five volumes dealing with herbal medicine, both maintained that the rhizome of common male fern would expel intestinal worms. English herbalist John Gerard (c. 1545-1612) writes: “The roots of the male fern, being taken in the weight of half an ounce, driveth forth long flat worms, as Dioscorides writeth, being drunke in mede or honied water, and more effectually if it be given with two scruples, or two third parts of a dram of scammonie, or of black hellebore: they that will use it, must first eat garlicke.”

In 1855, Swiss apothecary M. Peschier states that by digesting buds of common male fern in sulphuric ether, and using this tincture against tapeworms, they were killed with constant success. (Source: M. Peschier 1855. Bibliothèque Universelle de Genève, XXX, p. 205)

The pounded rhizome has the same effect. Due to the great toxicity of this drug, it is only used nowadays, when alternative drugs are inadequate. A decoction of the rhizome has also been used to treat certain fungal infections.

 

 

This common male fern has sprouted a very unusual place, in a crack in a wall, Copenhagen Central Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Fabaceae Pea family
Flowers of the subfamily Faboideae, to which the species below belong, have 5 petals, forming a unique structure. The upper petal, called the standard, is large and often reflexed, covering and protecting stamens and pistil. The two lateral petals, called the wings, are of equal size, surrounding the two bottom petals, which are free at the base, but fused at the tip, forming what is called the keel, as it resembles the keel of a boat. They enclose stamens and pistil.

 

Anthyllis Kidney-vetch
This genus contains somewhere between 22 and 32 species of herbs, some shrubby, distributed from Europe eastwards to the Ural Mountains, southwards to northern Africa, the Arabian Peninsula, and Iran, with an isolated occurrence in Ethiopian mountains.

The generic name is derived from Ancient Greek anthyllio, diminutive of anthos (‘flower’).

 

Anthyllis vulneraria Common kidney-vetch
A very diverse plant with numerous subspecies and intermediate forms, in Europe alone about 24 subspecies, some of which may be separate species.

This perennial, with ascending or upright stems, varies from dwarf plants 5 cm tall to shrubby herbs that may reach a height of 40 cm. Basal leaves often have only a terminal elliptic leaflet, to 8 cm long, lateral leaflets 1-4 pairs, much reduced in size, or often missing. Stem leaves have 2-7 pairs of leaflets, ovate, elliptic, or lanceolate, lateral ones to 2.5 cm long and 8 mm wide, terminal leaflet to 6 cm long and 2 cm wide, rounded.

The inflorescence is a compact flowerhead with numerous flowers, to 1.9 cm long, golden-yellow in most subspecies, occasionally whitish, orange, or red. The white-haired calyx is inflated, with uneven teeth. The pod is ovoid, hairy, dark, one-seeded.

This plant is distributed in the entire Europe, eastwards to the Ural Mountains, southwards to northern Africa and Iran, with an isolated occurrence in Ethiopian mountains, growing in dry grasslands and coastal dunes, along roads and embankments, and in other disturbed areas, preferably on calcareous soils, from the lowland up to alpine regions.

The specific name is derived from the Latin vulnus (‘wound’), alluding to the former usage of the plant for healing wounds. It was also used for cough.

 

 

Common kidney-vetch, subspecies carpathica, Copenhagen Central Station, Denmark. The grass is false oat-grass (Arrhenatherum elatius). (Photos copyright © by Kaj Halberg)

 

 

 

Lathyrus Vetchling
A large genus with about 180 species, found almost worldwide in temperate areas, apart from southern Africa, Southeast Asia, and Australasia. Most species are climbing herbs, with tendrils at the tip of the leaves, but some are erect shrubs. When the pods are mature, the two halves twist into spirals, hereby spreading the seeds a considerable distance.

The generic name is a Latinized version of Ancient Greek lathyros, the classical name of the chickling pea (L. sativus).

 

Lathyrus clymenum Spanish vetchling
This colourful plant is restricted to the Mediterranean area, found eastwards to Turkey and Jordan.

On the Greek island Santorini, the plant has been cultivated for more than 3,500 years. The seeds are used to prepare a special dish, called fava santorinis.

The specific name is derived from Ancient Greek klymenos (‘renowned’).

 

 

Spanish vetchling, growing along an abandoned railway, San Cataldo, Sicily. (Photo copyright © by Kaj Halberg)

 

 

 

Melilotus Melilot, sweet clover
About 20 species of herbs, distributed in temperate and subtropical areas of Eurasia and North Africa.

The leaves are trifoliate.

The generic name is derived from Ancient Greek meli (‘honey’) and lotus (see Lotus above). These plants are much visited by bees. The common name sweet clover refers to the fragrance of the flowers. They are also widely cultivated as nitrogen-producers.

 

Melilotus albus White melilot
Also known as white sweet clover, this plant is a native of southern and eastern Europe, eastwards to Central Asia. In former days, it was cultivated in other parts of Europe, and in North America, as a fertilizing plant, as its root nodules – like with other members of the pea family – contain bacteria, which are able to fixate nitrogen from the air. When these bacteria die (and they are very short-lived), the plant can utilize the nitrogen. When the plant dies, the nitrogen is released into the soil, for other plants to utilize. It also makes excellent hay.

Today, white melilot is not cultivated on a larger scale. Nevertheless, it has become widely naturalized in most temperate and subtropical areas of the world, especially in abandoned lots and along newly established roads.

The specific name is Latin, mening ‘white’.

 

 

White melilot, growing up a fence near Aarhus Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

In this picture, it grows near tracks at Alvesta Railway Station, southern Sweden, with sycamore maple (Acer pseudoplatanus) and common viper’s bugloss (Echium vulgare) in the foreground. (Photo copyright © by Kaj Halberg)

 

 

 

Trifolium Clover
A huge genus with about 300 species, found in temperate and subtropical areas of Eurasia, Africa, and America.

The pod is small, containing a single seed.

The generic name was the classical Latin word for clover, meaning ‘with 3 leaves’, referring to the trifoliate leaves of the genus.

 

Trifolium campestre Low hop clover
This plant, growing to 30 cm long/tall, has distinctive yellow flowerheads, which superficially resemble hop flowers, hence its common name. The finely serrated leaves have very conspicuous veins.

It is native from the British Isles eastwards to the Ural Mountains, and from northern Europe southwards to northern and eastern Africa, Arabia, Iran, and Afghanistan, growing in grasslands, fields, forest edges, wastelands, and along roads.

The specific name is Latin, meaning ‘growing in fields’.

 

 

Low hop clover, growing in gravel together with the red form of Herb Robert (Geranium robertianum), narrow-leaved plantain (Plantago lanceolata), red fescue (Festuca rubra), and wall barley (Hordeum murinum), Copenhagen Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Vicia Vetch
This genus with about 250 species is native to temperate and subtropical regions of Eurasia, Africa, and the Americas. In the Tropics, these plants are restricted to mountains.

The generic name is the classical Latin word for vetch.

 

Vicia tenuifolia Fine-leaved vetch
Stem erect, branched, to 1 m long, leaves pinnate, ending in a branched tendril and with 6-13 pairs of leaflets, which are linear or linear-lanceolate, 1.5-3 cm long and to 6 mm wide, smooth above, downy below, margin entire.

The inflorescence is a raceme with up to 30 flowers, usually 10-15 cm long, but sometimes to 30 cm, densely hairy. Corolla to 1.8 cm long, blue or purple, rarely white, but keel often white, standard to 1.5 cm long, keel a bit shorter. Flowering occurs June-September.

A widely distributed species, found in the major part of Europe and northern Asia, eastwards to central Siberia, southwards to Morocco, Jordan, the Himalaya, northern China, and Japan, growing at forest edges and in open areas.

The specific name is Latin, meaning ‘with thin leaves’.

 

 

Fine-leaved vetch, growing along an abandoned railway, San Cataldo, Sicily. (Photo copyright © by Kaj Halberg)

 

 

 

Geraniaceae Cranesbill family

 

Geranium Crane’s-bill
A large genus with about 380 species of herbs, mainly distributed in temperate areas, in the subtropics and tropics restricted to mountains.

Stipules (leaf-like appendices at the base of leaves) are often distinct on these plants. After flowering, the style forms a long, straight or up-curved beak, which separates into 5 elastic spring-like coils, each containing a single seed that is expelled, usually when the style is touched.

The generic name is derived from Ancient Greek geranos (‘crane’), alluding to the fruit, whose shape resembles a crane’s bill.

In German, these plants are called Storchschnabel (‘stork’s-bill’). In 1542, German physician and botanist Leonhart Fuchs (1501-1566) writes in his great herbal De Historia Stirpium Commentarii Insignes (‘Notable Commentaries on the History of Plants’): “Storckenschnäbel seind von den Griechen unnd Lateinischen genemt worden Gerania, darumb das sie am obersten teyl des stengels bringen ein köpfflin mit langen schnäbelin, nit anderst dann die Krench oder Storcken. Unnd daher kompt es, das zu unsern zeiten würdt Rostrum ciconiae geheyssen, das ist Storckenschnabel.” (‘By the Greeks and the Latinos, stork’s-bill is called Gerania, because at the top of the stem they bring forth a small head with long bills, no different from the bill of cranes or storks. And therefore, in our time, they are called Rostrum ciconiae, i.e. stork’s-bill.’)

In English, the name stork’s-bill refers to members of the genus Erodium, likewise of the crane’s-bill family.

 

Geranium purpureum Purple cranesbill
A widespread plant, native to western and southern Europe, northern Africa, and the Middle East, eastwards to Iran. It resembles the well-known Herb Robert (below), but has dark purple stems, and the lobes on the leaves are more rounded.

 

 

Purple cranesbill, growing along an abandoned railway, San Cataldo, Sicily. (Photos copyright © by Kaj Halberg)

 

 

 

Geranium pyrenaicum Hedgerow crane’s-bill, mountain crane’s-bill
Stem to 70 cm tall, hairy, glandular-hairy above. Leaves evergreen, long-stalked, to 7 cm across, rounded in outline, lower ones divided about halfway into 5-9 blunt segments with rounded teeth, upper ones more sharply divided. Stipules small, hairy, reddish-brown. The inflorescence is a loose raceme, flower stalks glandular-hairy, petals 5, purplish, violet, or pink, notched, to 1 cm long.

This plant grows in a variety of habitats, including open forests, meadows, grazing grounds, and wastelands. It is native to mountains of central and southern Europe and North Africa, eastwards to Ukraine, the Caucasus, and northern Iran.

It is widely cultivated and has become naturalized in numerous countries, northwards to Scandinavia, Finland, and European Russia, and also in north-eastern North America and Australia.

 

 

Hedgerow crane’s-bill, growing along an abandoned railroad track, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Geranium robertianum Herb Robert
This plant is native to most of Europe, and is also found in North Africa, western Asia, and parts of North America. It is very common in forests, but also grows in open habitats, including stony beaches and along railroad tracks. It is very variable, plants growing in shadowy places mostly having green leaves and reddish stems, whereas plants growing in open places most often have crimson stems and leaves.

In former days, this species was known as Saint Robert’s herb, like the specific name referring to French abbot and herbalist Robert de Molesme (c. 1028-1111), one of the founders of the Cistercian Order. He used it to cure people, suffering from various diseases, including diarrhea, liver and gall bladder problems, toothache, and wounds. American native tribes also utilized it medicinally.

In his booklet Chrut und Uchrut (in English called Weeds – a useful booklet on medicinal herbs), from 1911, Swiss priest and herbalist Johann Künzle (1857-1945) states the following: “The application of Herb Robin is also very effective against abscesses and inflammations of cattle. Glory be to God.”

 

 

Herb Robert with green leaves (top), and with both green and red leaves, observed at Copenhagen Railway Station, Denmark. (Photos copyright © by Kaj Halberg)

 

 

The red-leaved form of Herb Robert, growing abundantly between railroad tracks at Aarhus Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

These plants at Nyborg Railway Station, Denmark, have purple leaves. (Photo copyright © by Kaj Halberg)

 

 

 

Malvaceae Mallow family
Numerous members of the subfamily Malvoideae are described on the page Plants: Mallows and allies.

 

Malva Mallows
A genus with about 52 species, widespread in temperate, subtropical, and tropical regions of Eurasia and Africa.

The generic name is the classical Latin word for mallow, adapted from the classical Greek word for the same plants, malakhe, originally from Hebrew malluah, a plant used as salad. The common name is a corruption of Old English mealwe, an adaptation of the Latin name. The word for the colour mauve was adapted from the French name of these plants.

 

Malva sylvestris Common mallow
This species is native to temperate regions of Asia, Europe, and North Africa, but has been introduced elsewhere as an ornamental. It has become naturalized in many countries, including Australia, the United States, Canada, and Mexico.

It grows in open areas, including fallow fields, along hedges, and on the inner part of beaches. Stem to 1 m long, erect or ascending, leaves roundish, toothed, with 5-7 lobes, flowers in axillary clusters, petals pale violet or bright pinkish-purple with darker veins.

Several places around the Mediterranean, the leaves are steamed with garlic and tomatoes, eaten as an appetizer or a salad. In Egypt, the leaves are made into a stew-like vegetable dish, known as khobeiza. The leaves were also eaten as a vegetable in Europe in the 19th Century. The flowers were spread on doorways and woven into garlands for celebrating May Day.

In traditional medicine, the seeds were taken internally in tea to relieve irritation or inflammation of the mucous membranes, and the leaves were used in a poultices to soften the skin. A yellow dye can be obtained from the plant.

The specific name is Latin, meaning ‘growing in forests’ – not an appropriate name, as it grows in open areas.

 

 

Common mallow, Copenhagen Central Station, Denmark. The other plants are common mugwort (Artemisia vulgaris) and false oat-grass (Arrhenatherum elatius). (Photo copyright © by Kaj Halberg)

 

 

 

Oleaceae Olive family

 

Olea
This genus contains about 33 species of small trees, widely distributed in southern Europe, Africa, Madagascar, the Middle East, Arabia, subtropical and tropical Asia, and Australia.

The generic name is derived from Ancient Greek elaia, the classical name of the olive tree (below). It is the origin of the word ‘oil’.

 

Olea europaea Olive tree
The olive tree has been cultivated for more than 3,000 years. It is native to areas around the Mediterranean Sea, and is widely planted elsewhere in subtropical areas due to the excellent edible oil, extracted from the fruit flesh and seeds.

Its usage is described in depth on the page Plants: Plants in folklore and poetry.

 

 

This olive tree has sprouted along an abandoned railway, San Cataldo, Sicily. (Photo copyright © by Kaj Halberg)

 

 

 

Onagraceae Evening-primrose family

 

Chamaenerion Willow-herb
This genus with 8 species is widespread in montane and subarctic areas of the Northern Hemisphere, in Europe, North Africa, Asia, and North America. The fruit is a long capsule, splitting longitudinally, and the seeds have long hairs, an adaptation for wind-spreading.

There is much controversy as to the name of this genus. Initially, Chamaenerion may have originated as early as 1561. It is derived from Ancient Greek khamai (‘near the ground’) and nerion, the Ancient Greek name of the oleander (Nerium oleander), alluding to the oleander-like leaves of rosebay willow-herb (below). In 1753, Swedish botanist Carl Linnaeus (1707-1778) placed these plants in the genus Epilobium (below). However, many botanists disregarded his decision, preferring Chamaenerion.

In 1818, French naturalist Constantine Samuel Rafinesque-Schmaltz (1783-1840) proposed the name Chamerion, suggesting it as either a subgenus or genus. Rafinesque had his own peculiar rules of botanical nomenclature, regarding it as appropriate to shorten existing generic names. His new name, however, was not widely accepted until published in 1972 by Czech botanist Josef Ludwig Holub (1930-1999), who designated a different type species, Epilobium amenum. However, as this species is now included in C. angustifolium, Chamaenerion has precedence over Chamerion. Some authorities still include these plants in Epilobium. (Source: A.N. Sennikov 2011. Chamerion or Chamaenerion (Onagraceae)? The old story in new words, Taxon 60 (5): 1485-1488)

The common name alludes to the similarity of the leaves of rosebay willow-herb to those of certain species of willow (Salix).

 

Chamaenerion angustifolium Rosebay willow-herb, fireweed
The stem is erect, unbranched, smooth, very leafy, to 2.5 m tall, but often much lower. Leaves spirally arranged up the stem, stalkless, narrowly lanceolate, almost smooth, to 23 cm long and 3.5 cm broad, margin entire, lateral veins at a right angle from the mid-vein. Inflorescence is a lax, terminal, leafless raceme, to 50 cm long, flowers horizontal, sepals 4, linear, reddish-purple, to 2 cm long, petals 4, obovate, spreading, pink or magenta (rarely white), to 2.5 cm long and 1.5 cm broad. Style to 2 cm long, curved downwards, stigma deeply 4-lobed, stamens to 2 cm long, out-curved.

This gregarious herb often forms large growths, especially in open disturbed habitats, such as forest clearings and abandoned fields. An example of the latter is described on the page Nature Reserve Vorsø: Expanding wilderness.

It is very widely distributed in northern temperate and subarctic areas, southwards to Morocco, Pakistan, northern Indochina, Korea, and northern United States. In the Alps and the Pyrenees it grows up to elevations around 2,500 m, in the Himalaya up to about 4,700 m.

The specific name is Latin, meaning ‘narrow-leaved’. The name fireweed refers to the ability of this species to rapidly colonize areas, burned by fire. Incidentally, it was one of the first plants to appear when the eruption of Mt. St. Helens, Washington State, had devastated a huge area in 1980.

 

 

Rosebay willow-herb, Aarhus Railway Station (top), and Køge Railway Station, both in Denmark. False oat-grass (Arrhenatherum elatius) is also present in the upper picture. (Photos copyright © by Kaj Halberg)

 

 

 

Epilobium Willow-herb
About 200-220 species of herbs, distributed almost globally. Many species are glandular-hairy. The fruit is a long capsule, splitting longitudinally, and the seeds have long hairs, an adaptation for wind-spreading.

The generic name was used as early as 1561 for E. angustifolium (today Chamaenerion angustifolium, above) by Swiss physician and naturalist Conrad Gessner (1516-1565) in his unfinished work Historia plantarum. He composed the name from Ancient Greek ion epi lobion (‘violet on a pod’), lobion literally meaning ‘fruit of the cowpea (Vigna unguiculata), alluding to the similarity of the flower colour to that of certain violets (Viola), and to its pod-like fruit.

The common name refers to the similarity of the leaves of some species to those of certain species of willow (Salix).

 

Epilobium ciliatum Fringed willow-herb
This plant, also known by the names American willow-herb and Epilobium adenocaulon, is native to much of North America, southern South America, and East Asia, but has been accidentally introduced to many other places. It may grow to about 1.5 m tall, found in a variety of habitats, including waste plots, roadsides, and along railways.

The specific name is Latin, meaning ‘hairy’ or ‘fringed’, referring to the fine hairs on stems, leaves, and seed pods.

 

 

Fringed willow-herb, Aarhus Railway Station, Denmark. The small plant is procumbent pearlwort (Sagina procumbens, see above). (Photo copyright © by Kaj Halberg)

 

 

 

Epilobium hirsutum Great willow-herb
A robust, hairy plant, growing to 2 m tall, with leaves to 12 cm long and 3.5 cm wide. It perfers damp habitats, such as riversides and ditches, but may also be found in drier areas.

It is distributed in temperate regions, from Europe eastwards across Asia to China and Japan, and also in Arabia and large parts of Africa.

The specific name is Latin, meaning ‘hairy’, alluding to the dense layer of hairs on the stem.

 

 

Sjælland 2012-16
Great willow-herb, growing among railroad tracks at Køge Railway Station, Zealand, Denmark, an unusually dry habitat of this species. (Photo copyright © by Kaj Halberg)

 

 

 

Epilobium montanum Broad-leaved willow-herb
This common species, which grows to 60 cm tall, is easily identified by its smooth, serrated, ovate leaves. It is distributed from Europe eastwards to central Siberia, southwards to the Mediterranean and Iran. It has also been introduced elsewhere, including North America, Japan, and New Zealand. It grows in open forests and shrubberies, and also in open areas, including gardens.

The specific name is Latin, meaning ‘found in mountains’ – an odd name, as it is also common in the lowlands.

 

 

Broad-leaved willow-herb, growing along railroad tracks, Aarhus Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Oenothera Evening-primrose
This large genus, comprising about 145 species, is native to the Americas, but many species are widely cultivated for their attractive flowers, which unfold late in the evening, stay open at night, and wither during the following day. This is an adaptation to being pollinated by insects that are active during the night, including moths.

These plants vary greatly in size, from dwarves 10 cm high to giants 3 m tall. Most species have yellow flowers, some white, purple, pink, or red. Despite their name, they are not closely related to the true primroses (Primula).

The meaning of the generic name is uncertain.

 

Oenothera glazioviana Large-flowered evening-primrose
This impressive plant, previously known as O. erythrosepala, may grow to 2 m tall, leaves stalked, alternate, lanceolate, strongly crinkled, to 15 cm long.

It is a native of Brazil, but is widely cultivated as an ornamental plant, often becoming naturalized.

The specific name honours French botanist and landscape designer Auguste François Marie Glaziou (1828-1906). In 1858, Emperor Dom Pedro of Brazil (1825-1891), whose real name was no less than Pedro de Alcântara João Carlos Leopoldo Salvador Bibiano Francisco Xavier de Paula Leocádio Miguel Gabriel Rafael Gonzaga, requested Glaziou to become director of parks and gardens in Rio de Janeiro. While in Brazil, Glaziou also collected numerous plants.

 

 

Large-flowered evening-primrose, Ringsted Railway Station, Zealand, Denmark. Common viper’s bugloss (Echium vulgare) and horsetail fleabane (Erigeron canadensis) are growing in front of the wagon. (Photo copyright © by Kaj Halberg)

 

 

 

Papaveraceae Poppy family

 

Papaver True poppies
This genus contains about 150 species, found in subarctic, temperate, and subtropical regions of the Northern Hemisphere, with a single species in southern Africa.

The generic name is the classical Latin term for poppies.

 

Papaver argemone Prickly-headed poppy
This delicate plant may be told from other red poppies by having stiff hairs on stem and fruit, and it often has black blotches at the base of the petals. It is native to temperate areas of Europe, eastwards to Ukraine and the Caucasus, and in North Africa.

The specific name derived from Ancient Greek argemos, meaning ‘a white spot on the eye’ (cataract). According to the Ancient Greeks, a species of Argemone, likewise in the poppy family, was able to cure cataract. Why the name was applied to prickly-headed poppy is not clear.

 

 

Prickly-headed poppy, growing at Nyborg Railway Station, Funen, Denmark. To the left withering stems of thale cress (Arabidopsis thaliana). (Photo copyright © by Kaj Halberg)

 

 

 

Papaver dubium Long-headed poppy
This plant is very common in the major part of Europe, northern Africa, and western Asia, eastwards to the Himalaya. It mostly grows in fallow fields and waste places, and along newly established roads.

It is quite variable, growing to about 60 cm tall, stem with stiff bristles, basal leaves forming a rosette, stalked, hairy, to 15 cm long and 3 cm wide, pinnately lobed, each lobe with a fine spine at the end, stem leaves alternate, smaller, sessile. The showy flowers, sometimes to 7 cm across, have 4 orange-red petals. The similar corn poppy (P. rhoeas) has darker red petals, usually with a black spot at the base, which lacks in the long-headed poppy.

The specific name is Latin, meaning ‘in doubt’. Perhaps Swedish naturalist Carl Linnaeus (1707-78) was in doubt, when he named the plant in 1753, whether it was the same species as the corn poppy.

 

 

Long-headed poppy, Copenhagen Central Station, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Plantaginaceae Plantain family

 

Linaria Toadflax
A widespread genus, comprising about 185 species, distributed in the entire subarctic and temperate regions of the Old World. Many species have also become naturalized elsewhere, especially in North America. These plants are characterized by their flowers, which have a gaping mouth and a long spur.

The generic name is derived from the Latin linum (‘linen’ or ‘flax’), thus ‘resembling flax’, alluding to the leaves of some species, which superficially resemble those of flax.

The toad part of the common name has various explanations. In his Herball (1597), English herbalist John Gerard (c. 1545-1612) says that the name was applied to these plants, because the flowers “be yellow, having a mouth unto a frog’s mouth.” Some 60 years later, English herbalist William Coles (1626-1662), in his work Adam in Eden, or the Paradise of Plants, claims that the name arose “because toads will sometimes shelter themselves amongst the branches of it.”

 

Linaria repens Pale toadflax
Also known as creeping toadflax or striped toadflax, this species is native to south-western Europe, but is widely naturalized elsewhere in Europe, and in North America, presumably brought with soil, which acted as ballast on board cargo ships.

The specific name is Latin, meaning ‘creeping’, which is misleading, as the plant is often erect. The common name striped toadflax refers to the purple stripes on the whitish flowers.

 

 

The preferred habitat of pale toadflax is dry soils, and it is quite common along railroad tracks, here at Alvesta, southern Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Poaceae Grass family
Numerous grass species and their usage are described on the page Plants: Grasses.

 

Arrhenatherum
A small genus with 7 species, distributed from the entire Europe, including Iceland, eastwards to the Ural Mountains and Kazakhstan, southwards to northern Africa, Jordan, and Iran.

The generic name is derived from Ancient Greek arrhenos (‘male’) and atheros (‘bristle’), alluding to the fact that only the male flowers have awns.

 

Arrhenatherum elatius False oat-grass
This grass, occasionally growing to 1.8 m tall, has long, slightly hairy, rough leaves, to 1 cm broad. The panicle is up to 30 cm long, spikelets with angled, projecting awns, to 1.7 cm long, green or purplish. The panicles often remain into winter.

It is found from Iceland eastwards to the Ural Mountains, southwards to North Africa, Iran, and Kyrgyzstan. It is very common, growing in a wide variety of open habitats, for instance along roads, hedges, fields, and streams. It has been greatly favoured by the increased nitrogen fallout in later years.

The specific name is Latin, meaning ‘tall’. The common name alludes to its superficial resemblance to the cultivated oat (Avena sativa), described on the page Plants: Grasses.

 

 

Large growths of false oat-grass along an abandaned railroad track, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

False oat-grass, Fredericia Railway Station, Jutland (top), and Copenhagen Central Station, both Denmark. (Photos copyright © by Kaj Halberg)

 

 

Stems of false oat-grass, moving in the wind, Nyborg Railway Station. (Photo copyright © by Kaj Halberg)

 

 

Withered stems of false oat-grass along an abandaned railroad track, Nyborg. (Photo copyright © by Kaj Halberg)

 

 

 

Bromus Brome
A large genus with about 160 species, widely distributed on all continents, except Antarctica, and it is also absent from certain rainforest and desert areas.

In Ancient Greece, the generic name was applied to a grassy weed, presumably wild oat (Avena fatua). The word bromos means ‘noise’, alluding to the panicles rattling in the wind. Why the name was applied to bromes is not clear.

 

Bromus sterilis Barren brome
This grass, also called poverty brome or sterile brome, is very common, native to the major part of Europe, North Africa, and south-western Asia, eastwards to Kazakhstan and Kyrgyzstan. It grows in a wide variety of open habitats, including waste areas, roadsides, and gardens.

The specific and common names are derogatory, probably indicating that this species is worthless in all respects.

 

 

Barren brome, Nyborg, Denmark, at the railway station (top), and along an abandaned railroad track. (Photos copyright © by Kaj Halberg)

 

 

 

Hordeum Barley
This genus contains about 34 species, native to temperate and subtropical areas of the Northern Hemisphere, and also to South America and southern Africa.

The generic name is classical Latin term for the cultivated barley (H. vulgare), derived from Proto-Italic horzdeom (‘bristly’), alluding to the long, prickly awns of these plants.

 

Hordeum murinum Wall barley
Wall barley, also called false barley, is native to central and southern Europe, eastwards to the Ural Mountains and Kazakhstan, southwards to northern Africa, Arabia, and north-western Himalaya. It grows along roads, as a weed in fields, and in waste plots. It has readily adapted to a life in cities.

In China and Taiwan, the seeds are an ingredient in a green type of sticky rice dumplings, called qingtuan, which are consumed during the Qing Ming Festival (‘tomb-sweeping’). This festival is described on the page Culture: Graves.

The specific name is derived from the Latin murus (‘wall’), alluding to the fact that this plant often grows along stone fences and house walls.

 

 

Wall barley, growing along an abandoned railroad track, Nyborg, Denmark, together with dandelion (Taraxacum officinale), hedgerow crane’s-bill (Geranium pyrenaicum), and common evening-primrose (Oenothera biennis). (Photo copyright © by Kaj Halberg)

 

 

Here, wall barley is growing together with Herb Robert (Geranium robertianum), Copenhagen Central Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Poa
This huge genus, containing about 575 species, is cosmopolitan, only being absent from the polar regions and certain rainforest and desert areas.

The generic name is Ancient Greek, simply meaning ‘fodder’.

 

Poa annua Annual meadow-grass
This grass, in America commonly known as annual bluegrass, is native to a huge area, found from western Europe and northern Africa across Temperate Asia to the Pacific. It has also been spread to most other temperate and subtropical areas of the world, deliberately as a fodder plant, or accidentally. According to some authorities, it may have originated as a hybrid between Poa supina and Poa infirma.

It is a small plant, growing to a maximum of 25 cm tall, but often much lower. It is a very common weed in trampled places, such as trails and in cities.

 

 

Annual meadow-grass, Copenhagen Central Station, Denmark. The other plants are horsetail fleabane (Erigeron canadensis) and common sow-thistle (Sonchus oleraceus). (Photo copyright © by Kaj Halberg)

 

 

 

Polygonaceae Buckwheat family

 

Rumex Dock, sorrel
A large genus with about 200 species of herbs, some rather shrubby, with an almost worldwide distribution, and introduced species are often found in the few places where the genus is not native. Some are invasive, others are cultivated for their edible leaves.

The fruit is mostly a triangular nutlet, often winged, arranged in dense clusters, often conspicuously reddish-brown when ripe.

The generic name is the classical Latin word for sorrel. Dock is an old term applied to various plants with large leaves, whereas sorrel is a brownish-orange to pale chestnut colour, referring to the fruits of this genus.

 

Rumex acetosa Common sorrel
This plant occurs from Iceland and the British Isles eastwards to central Siberia, southwards to the Mediterranean, Iran, China, and Japan. It has also been introduced to many other places, including Australia, New Zealand, and North America.

Common sorrel has been cultivated for centuries. The leaves are full of vitamin C. They are edible when young, and are often used in salads, and applied to soups, sauces, or curries.

The common name sorrel is an old word for a brownish-orange to pale chestnut colour, referring to the brown fruits of the genus.

 

 

Common sorrel, Copenhagen Central Station, Denmark. The other plants are common mugwort (Artemisia vulgaris), cat’s-ear (Hypochaeris radicata), and red fescue (Festuca rubra). (Photo copyright © by Kaj Halberg)

 

 

 

Ranunculaceae Buttercup family

 

Clematis Clematis
This almost worldwide genus contains about 400 species of climbers, usually with a woody stem. The leaves are pinnate or trifoliate, and the fruit is a globular head of numerous nutlets, often with long silky hairs on the persistent styles.

These plants climb due to the sensitivity of the leafstalks which, when touching another plant or something else, start twisting, curling around the object.

The generic name is derived from Ancient Greek klema (‘climber’).

 

Clematis vitalba Traveller’s joy
A most vigorous climber, occasionally reaching a length of more than 20 m, with thick, woody stems to 6 cm in diameter. Leaves opposite, pinnate, with 5 (rarely 3) widely spaced leaflets, thin, sparsely hairy, entire or toothed. Flowers in umbel-like clusters, creamy-white, fragrant, to 3 cm across.

This pioneer plant grows in open forests, shrubberies, and fallow lands, especially on nitrogen-rich soils. It is distributed in central and southern Europe, North Africa, and the Middle East, eastwards to Afghanistan. Elsewhere, it is widely cultivated, often escaping to form naturalized growths. In New Zealand, it has been declared an unwanted organism. In the Alps and the Pyrenees, it may be encountered up to elevations around 1,300 m.

It contains the toxic protoanemonin, which may cause reddening and itching of the skin. In the past, beggars smeared juice of the leaves on the skin to cause ulcerations, which might arouse pity in people. In Tuscany, spring sprouts were formerly used in a certain kind of omelettes, called vitalbini, which was maybe not so wise due to its toxicity.

Baskets were produced from the tough stems. In the Alps, in former days, children would smoke dry stems as cigarettes.

The specific name is the Italian name of the plant. In England, the name traveller’s joy was given in allusion to the profusion of pleasant feathery seedheads of this plant in the dark months leading up to Christmas. The name old man’s beard also refers to the seedheads. An old German folk name was Teufelzwirn (‘devil’s twine’), referring to its toxicity.

 

 

Traveller’s joy, growing over an abandoned railway, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Rosaceae Rose family

 

Geum Avens
A genus with about 50 species, widespread in Eurasia, Africa, New Zealand, and the Americas. Most species have yellow flowers, some red, orange, or white. A gorgeous orange species is shown on the page Travel episodes – Chile 2011: The white forest.

At maturity, a silky tuft of brownish hairs grows from the styles, which has given rise to a popular German name of these plants, Petersbart (‘Peter’s beard’), probably referring to St. Peter.

German Dominican friar, philosopher, scientist, and bishop Albertus Magnus (c. 1200-1280) says: “Wo diss wurtz in dem huse ist, da mag der tufel nicht schaffen, und fluhet sie, und darumb ist sie gebenedeyt für alle anderen wurtzeln.” (‘If you have this herb in your house, the Devil does not want to be there and flees from it, and this is why this herb is better than all other herbs’).

The generic name is the classical Latin name for avens. The English name is derived from the Latin avencia, which was the name of a kind of clover. Why it was applied to these plants is not known.

 

Geum urbanum Wood avens
This herb thrives in shady forests, but may also be encountered along hedges, fences, and house walls, etc. It is found throughout Europe, eastwards to the Ural Mountains, Kazakhstan, the Hindu Kush and the Alborz Mountains of Iran, and also in north-western Africa.

At an early point, it was associated with Christianity due to its leaves growing in threes and its petals in fives – the threes reminiscent of the Holy Trinity (the Father, the Son, and the Holy Spirit), the fives with the Five Sacred Wounds that Jesus suffered on the Cross (four from the nails in hands and feet, and one from the spear that penetrated his side).

In folklore, wood avens was credited with the power to drive away evil spirits, and to protect against rabid dogs and venomous snakes. The root, which contains eugenol, were used as a spice in soups and also to flavour ale.

The specific name is Latin, meaning ‘grows in cities’ – not an appropriate name, as it is most common in forests.

 

 

Wood avens, growing at an abandoned railway, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Rubus Bramble, raspberry, and allies
This huge genus, comprising more than 1,400 species of prickly shrubs or creeping shrublets, is found all over the globe, except in Antarctica and certain deserts and tropical forests.

The fruit is highly distinctive, a globular head on the domed tip of the flower-stalk, consisting of fleshy carpels, usually with many nutlets on the surface.

The generic name is the classical Latin word for bramble (below).

 

Rubus caesius Dewberry
This species has a very wide distribution, from Scandinavia southwards to the Mediterranean, and from Ireland across Europe and western Asia to Xinjiang. It has also become naturalized in various other countries, including Canada, the United States, and Argentina. Its autumn foliage is a lovely crimson or wine-red, shown on the page Autumn.

The specific name is Latin, meaning ‘blue-grey’, alluding to the colour of the fruits.

 

 

Creeping stem of dewberry, Nyborg Railway Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Rubus fruticosus Bramble, blackberry
This species, also known as R. plicatus, is very variable, especially as regards the size and number of thorns on the stem, and the leaf shape. It is widely distributed in northern and central Europe, but has been introduced to many other countries.

It is highly invasive in some areas, forming dense thickets, which expel native vegetation and often threaten entire ecosystems. It is considered a noxious weed in many countries, including Australia, New Zealand, and the United States.

The specific name is Latin, meaning ‘shrubby’.

Other pictures, depicting this species, are shown on the page Autumn.

 

 

Bramble, growing over an abandoned railway, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Rubiaceae Bedstraw family

 

Galium Bedstraw
This huge genus, comprising about 640 species, is native to almost the entire world, with the exception of Antarctica and some rainforest and desert areas.

The generic name is derived from Ancient Greek gala (’milk’), alluding to the usage of yellow bedstraw (G. verum) in rennet. The common name refers to the usage of this plant in beds. More about these issues on the page Plants: Plants in folklore and poetry.

 

Galium aparine Cleavers
The specific name of this scrambling plant is derived from Ancient Greek apairo (‘to seize’) – a most descriptive name for this species. Not only the seeds are armed with hooks, but also stems and leaves, which aid the plant in climbing or scrambling over other plants.

In his delightful book All about Weeds, American botanist Edwin Spencer (1881-1964) has the following comment about this plant: “When one walks into some loose, trailing weeds, and they all seem glad to be torn from their roots in order to ride away clinging to his trousers, he has made contact with one of the many bedstraws (…) but if the stems cling to the clothing, the chances are that the find is one of two of them, Galium aparine or Galium asprillum. These are the two with the rough stems – the two that have this unique way of scattering their seeds. For of course those straws cling to the sheep’s wool and the dog’s hair just as they do to clothing, and sometimes they and their seeds are carried a long way by animals.”

Cleavers is also known by many other names, which allude to is hooked seeds and its scrambling habit, including catchweed, sticky bob, robin-run-the-hedge, and sticky willy.

This species is native to the major part of Europe, North Africa, and Temperate Asia, eastwards to Japan. It may also be indigenous in North America, and has become naturalized in Mexico, Central America, South America, Australia, New Zealand, and parts of Africa.

 

 

Cleavers, growing at Nyborg Railway Station, Funen, Denmark. Withering stems of thale cress (Arabidopsis thaliana) are also present. (Photo copyright © by Kaj Halberg)

 

 

 

Sapindaceae Soapberry family

 

Acer Maple
In former days, maples constituted a separate family, Aceraceae, which is now regarded as belonging to Hippocastanoideae, a subfamily of the soapberry family.

These plants, comprising about 130 species of deciduous trees or large shrubs, are widespread in the Northern Hemisphere, with most species in Asia, others in Europe, northern Africa, and North and Central America. Only a single species occurs in the Southern Hemisphere.

The leaves are palmate in most species, and the foliage often turns a brilliant red or yellow in autumn. A number of pictures, depicting this autumn foliage, are found on the page Autumn.

The fruit consists of 2 connected single-seeded units, each with a long wing. When ripe, the fruits are often propelled a considerable distance by the wind. An account of the effectiveness of this spreading is described on the page Nature reserve Vorsø: Expanding wilderness.

The generic name is the classical Latin word for maples.

 

Acer platanoides Norway maple
This maple is native to Europe, found from southern Scandinavia southwards to the Pyrenees, Italy, and the Balkans, eastwards to Ukraine, and thence southwards to the Caucasus and Turkey.

At an early stage, it was introduced to North America, where it is now regarded as an invasive plant in many eastern states. For this reason, Massachusetts and New Hampshire have banned planting of this tree.

In autumn, the foliage attains a bright yellow colour.

The specific name is Latin, meaning ‘resembling Platanus‘ (plane trees), referring to the angular leaf lobes, which resemble those on the leaves of the oriental plane tree (Platanus orientalis).

 

 

Fallen leaves of Norway maple on an abandoned railway, Nyborg, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Acer pseudoplatanus Sycamore maple
This species, which is a native of Central Europe, was introduced to Britain around 1500. It has also become naturalized in other parts of Europe, and in Australia, New Zealand, and North America. In many places, it has become invasive, easily spreading by its winged seeds, which are produced in the tens of thousands on a single large tree.

An example of the effective spreading of sycamore maple is seen in nature reserve Vorsø, Horsens Fjord, Denmark, where former fields were abandoned in two steps, in 1928 and in 1978. In both cases, thousands of maple seeds, stemming from a few trees in plantations at the edge of the fields, were spread by the wind. The succession of these maple forests is described in detail on the page Nature Reserve Vorsø: Expanding wilderness.

The specific name is Latin, meaning ‘the false Platanus‘ (plane trees), referring to the leaves resembling the leaves of these trees.

 

 

Sycamore maple, growing at Nyborg Railway Station, Funen, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Solanaceae Nightshade family

 

Solanum Nightshade
This is one of the largest plant genera in the world, counting more than 1,300 species, including a number of very important food plants, such as potato (S. tuberosum), tomato (S. lycopersicum), tamarillo (S. betaceum), and eggplant (S. melongena).

The generic name is of unknown origin, possibly derived from the Latin sol (’sun’) referring to the fact that plants of this genus prefer to grow in sunny places. The popular name is a corruption of an ancient Germanic name of these plants, Nachtschatten, of unknown meaning. As Schatten also means ‘shadow’ in German, the name nightshade was introduced, but apparently this is a misnomer.

 

Solanum dulcamara Bittersweet nightshade
This proliferous climber is very common in the major part of Europe, in North Africa, and in a belt across the central parts of Asia, eastwards to Japan, and it is also commonly naturalized in North America, where it is considered an invasive weed. It is partial to humid areas, where it often forms dense, almost impenetrable growths.

Previously, it was much used in traditional medicine for treatment of a large number of ailments, including skin diseases, warts, tumours, arthritis, rheumatism, bronchitis, heart ailments, jaundice, cough, diarrhoea, back and joint pains, and eye inflammations. The fruits, marinated in vinegar, and also a decoction of the root, were applied to cancerous sores and other swellings. It was also used as a diuretic, a purgative, and an emetic.

However, the plant is very poisonous and might paralyze the central nervous system, slow down heartbeat and respiration, and lower the temperature, causing vertigo, delirium, convulsions, and sometimes even death.

The specific name is Latin, mesning ‘bittersweet’, alluding to the taste of leaves and berries, being both bitter and sweet.

 

 

Bittersweet nightshade prefers to grow in humid areas, sometimes forming dense, almost impenetrable growths. However, it is also able to thrive in drier areas, such as this plant, growing near a railroad track at Horsens Station, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

(Uploaded July 2018)

 

(Latest update August 2025)