Portraits

 

 

This bull black rhino (Diceros bicornis) is sniffing a tuft of grass, into which a female has urinated, baring its lips in a posture, called flehmen. The inhaled air passes a special sensing organ, which is able to detect whether the female is in heat. – Ngorongoro Crater, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

Courting pair of Australian gannet, Muriwai Beach, New Zealand. (Photo copyright © by Kaj Halberg)

 

 

This excited flap-necked chameleon (Chamaeleo dilepis), with expanded gular sack, displays numerous black spots, Masasi, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

Female dromedaries (Camelus dromedarius) are often affectionate towards each other. – Thar Desert, Rajasthan, India. (Photo copyright © by Kaj Halberg)

 

 

 

Mammals

 

Atelidae
This is a family of New World monkeys, comprising about 26 species, placed in four genera: Ateles (spider monkeys), Alouatta (howler monkeys), Brachyteles (woolly spider monkeys), and Lagothrix (woolly monkeys).

 

Ateles Spider monkeys
This genus of 7 species is found from southern Mexico southwards to Brazil, living in the upper stratum of tropical forests. These monkeys are characterized by their disproportionately long limbs, which have given them their name, and their long, prehensile tail, which is used as a fifth limb. They live in bands, comprising up to 35 members, which will disperse during the day to feed.

All species of spider monkeys are threatened due to habitat destruction and hunting for food, and two species, the black-headed spider monkey (A. fusciceps) and the brown spider monkey (A. hybridus), are critically endangered.

 

Ateles geoffroyi Geoffroy’s spider monkey
Six subspecies of this monkey, which is also called black-handed or Central American spider monkey, are distributed from south-eastern Mexico eastwards to Panama. It mainly occurs in evergreen rainforest, but may also be found in deciduous forest. The species is threatened due to habitat loss, which has been severe across its entire range. It is estimated that it has declined by as much as 50% during the last 50 years. Today, it mainly survives in protected areas.

 

 

This red-bellied spider monkey, ssp. frontatus, also called black-browed spider monkey, is feeding on coffee-like fruits, Tortuguero National Park, Limón, Costa Rica. (Photo copyright © by Kaj Halberg)

 

 

 

Bovidae Cattle etc.
This large family, comprising about 47 genera and c. 143 species, are cloven-hoofed, ruminant animals, including cattle, antelopes, sheep, goats, and many others.

 

Aepyceros melampus Impala
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

Male impala, Tarangire National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Antidorcas marsupialis Springbok
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

Springbok, Kalahari Gemsbok National Park, South Africa. (Photo copyright © by Kaj Halberg)

 

 

 

Bos bison American bison
The sad fate of this animal is described on the page Folly of Man.

 

 

American bison bull, rubbing on a fence, Badlands National Park, South Dakota, United States. (Photo copyright © by Kaj Halberg)

 

 

 

Bos grunniens Yak
The yak is a high-altitude species, which used to roam the Central Asian highlands in large numbers. It is adapted to a life in this harsh environment, having a luxurious fur, which keeps it warm in temperatures below -30o Centigrade. – A Nepalese legend, which explains how the yak got its rich fur, is related on the page Animals as servants of man: Water buffalo.

The yak was domesticated by nomadic tribes as early as c. 5000 B.C., and today the population is estimated at 14 million, the vast majority in Chinese territories. The population of wild yak may be fewer than 15,000, and though it is legally protected, illegal hunting still takes place and may threaten this magnificent animal with extinction.

The scientific name is Latin for ‘grunting ox’ – a most descriptive name, as it grunts incessantly.

 

 

Yak, resting in front of a barberry bush, Dingboche, Khumbu, eastern Nepal. (Photo copyright © by Kaj Halberg)

 

 

 

Bos taurus Cattle
Catte, including the zebu ox, are described on the page Animals: Animals as servants of Man.

 

 

Inquisitive zebu ox, Pushkar, Rajasthan, India. (Photo copyright © by Kaj Halberg)

 

 

Zebu ox, dark morph, Jaisalmer, Rajasthan. (Photo copyright © by Kaj Halberg)

 

 

Longhorned Ankole cattle, an indigenous cattle breed of sub-Saharan Africa, Mubende, Uganda. (Photo copyright © by Kaj Halberg)

 

 

 

The Jersey is a British breed of small dairy cattle, originating from Jersey, one of the Channel Islands. Selective breeding has caused the milk production of these cows to increase to an average of 6,024 litres per year in the U.K., with some individuals yielding around 9,000 litres. The milk has a characteristic yellowish tinge and is high in butterfat (5.4%) and protein (3.8%).

As the Jersey adapts well to cold as well as hot climates, it has been exported to many countries around the world. Some countries have developed separate breeds.

 

 

Curious Jersey heifer, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

As its name implies, the Scottish Highland cattle originated in the Scottish Highlands, or maybe in the Outer Hebrides, first mentioned in the 6th Century A.D. In Scots, it is called Heilan coo, a name of Norse origin, meaning ‘highland cow’.

This breed is characterized by the long horns and the wavy coat, which comes in a variety of colours, including red, ginger, black, dun, yellow, white, or grey. The long coat allows it to spend the harsh Scottish winter outdoors. It is raised primarily for the meat, which is prized for its low content of cholesterol, and also for the milk, which generally has a very high butterfat content.

Today, Scottish Highland cattle is found in many countries around the world.

 

 

Scottish Highland cattle, Öland, Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Capra ibex Alpine ibex
At one point, this magnificent animal was almost hunted to extinction, only surviving in a few pockets in northern Italy. Due to the alarming decrease of the population, Victor Emmanuel, later to become king of Italy, declared the Royal Hunting Reserve of Gran Paradiso in 1856, and a protective guard was created for the ibex. In 1920, King Victor Emmanuel III donated the original 21 square kilometres to the country, and it became Italy’s first national park in 1922. Despite the park, ibex were poached until 1945, when only 419 remained. Since then, the population has increased, and there are now almost 4,000 in the park. It has been reintroduced to numerous other areas in the Alps, and also to Bulgaria and Slovenia.

In summer, the alpine ibex lives in rocky areas just below the snow line, at elevations between 1,800 and 3,300 m, descending to lower altitudes in the winter.

 

 

Male Alpine ibex, marked with an ear tag, Gran Paradiso National Park, Italy. (Photo copyright © by Kaj Halberg)

 

 

 

Connochaetes taurinus Blue wildebeest
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

White-bearded wildebeest, subspecies mearnsi, Ngorongoro Crater, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

During the annual wildebeest migration, these white-bearded wildebeest quench their thirst in the Grumeti River, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Kobus ellipsiprymnus Common waterbuck, defassa waterbuck
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

Male common waterbuck, ssp. ellipsiprymnus, chewing the cud, Hwange National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 


Male East African defassa waterbuck, ssp. harnieri, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Kobus kob Kob
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

Male Uganda kob, subspecies thomasi, resting in tall grass, Queen Elizabeth National Park, Uganda. (Photo copyright © by Kaj Halberg)

 

 

 

Madoqua kirkii Kirk’s dik-dik
Four subspecies of this diminutive antelope are distributed from extreme eastern Uganda, northern Kenya, and southern Somalia southwards to southern Tanzania, whereas a fifth, ssp. damarensis, which forms a disjunct population in western Angola and north-western Namibia, may constitute a separate species.

 

 

Kirk’s dik-dik, subspecies cavendishi, eating from a bush, Arusha National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Nilgiritragus hylocrius Nilgiri tahr
A rare species of sheep, which is restricted to mountains of South India, counting only around 3,100 individuals. Formerly, it was placed in the genus Hemitragus, together with the Himalayan tahr (H. jemlahicus) and the Arabian tahr (today called Arabitragus jayakari). However, recent genetic research has shown that it is more closely related to sheep of the genus Ovis than to tahrs, and, consequently, it was moved to a separate genus.

Several pictures, depicting this species, are shown on the page Quotes on Nature.

 

 

Eravikulam National Park, Kerala, where this picture was taken, is a stronghold of the Nilgiri tahr, housing an estimated 700-800 individuals, app. one-fourth of the total population. (Photo copyright © by Kaj Halberg)

 

 

 

Oreamnos americanus Mountain goat
Despite its name, this animal is not a true goat of the genus Capra, but is more closely related to serows (Capricornis), gorals (Naemorhedus), and the chamois (Rupicapra), sometimes referred to as goat-antelopes. It is endemic to mountainous areas of western North America, from southern Alaska southwards through western Canada to Oregon, northern Nevada, Utah, and Colorado.

The generic name is derived from Ancient Greek oros (‘mountain’) and amnos (‘lamb’), thus ‘mountain lamb’, presumably alluding to the white fur of the animal.

 

 

Confiding female mountain goat, Mount Rushmore, Black Hills, South Dakota. (Photo copyright © by Kaj Halberg)

 

 

 

Ovis canadensis Bighorn sheep
Three subspecies of this impressive animal are widely distributed in mountains of western North America, from British Columbia and western Alberta southwards through the Rocky Mountains and the Sierra Nevada to Baja California, north-western Mexico, and south-western Texas, as far east as North and South Dakota. This species was once very numerous, but the population has been much reduced by overhunting and introduction of diseases from livestock.

The desert bighorn sheep, ssp. nelsoni, occurs throughout the desert regions of the south-western United States and north-western Mexico.

 

 

Ram of desert bighorn sheep, ssp. nelsoni, Tucson Desert Zoo, Arizona. (Photo copyright © by Kaj Halberg)

 

 

 

Raphicerus campestris Steenbok
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

Female steenbok, Etosha National Park, Namibia. (Photo copyright © by Kaj Halberg)

 

 

 

Tragelaphus scriptus Bushbuck
This antelope is described on the page Animals – Mammals: Antelopes.

 

 

Female bushbuck, Victoria Falls National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Camelidae Camels
The origin of camels, and their relationship with people, is described on the page Animals: Animals as servants of Man.

 

Camelus dromedarius Dromedary, one-humped camel
This species is extinct in the wild, but is widely distributed as a domestic animal, from India across the Middle East to the Arabian Peninsula, Egypt, Somalia, and northern Kenya. It was probably first domesticated in Somalia or southern Arabia, around 3000 B.C.

The dromedary is described in detail on the page Animals: Animals as servants of Man.

 

 

 

Portrait of a dromedary, Jaisalmer, Rajasthan, India. (Photo copyright © by Kaj Halberg)

 

 

Decorated dromedary at a camel festival, Bikaner, Rajasthan. (Photo copyright © by Kaj Halberg)

 

 

Dromedaries, chewing their cud, Thar Desert, Rajasthan (top), and Sousse, Tunisia. (Photos copyright © by Kaj Halberg)

 

 

When a dromedary bull is in heat, froth is oozing out of his mouth. – Bikaner, Rajasthan. (Photo copyright © by Kaj Halberg)

 

 

 

Canidae Dog family
This family is described in depth on the page Animals – Mammals: Dog family.

 

Canis lupus ssp. familiaris Domestic dog
The domestication of the dog and its long association with Man is described in depth on the page Animals: Animals as servants of Man.

 

 

This dog in Guiyang, Guizhou Province, China, may have some Tibetan spaniel genes, and probably also some Pekingese, due to its very prominent lower jaw. (Photo copyright © by Kaj Halberg)

 

 

 

I asked the owner of this little terrier in Taichung, Taiwan, why it was wearing sunglasses. She said that it was to prevent the dog from getting cataract. (Photo copyright © by Kaj Halberg)

 

 

 

The English Springer Spaniel is a hunting dog breed, descended from Norfolk or Shropshire Spaniels in the mid-1800s. It was traditionally used for flushing out game, and for retrieving it. Today, it is also a popular family dog.

During a research trip to the Chukotka Peninsula, north-eastern Siberia, my companions and I visited the staff of a light house, situated at the tip of Kosa Ruskaya Koshka (‘Russian Cat’s Sandspit’). One of the dogs belonging to the staff was an English springer spaniel. Our trip to this area is related in detail on the page Travel episodes – Siberia 2011: Caterpillar trip in Chukotka.

 

 

(Photo copyright © by Kaj Halberg)

 

 

 

The short-legged dachshund was developed to chase foxes and badgers out of their dens, for the hunter to shoot them. In America, they have also been used to chase prairie dogs out of their dens. This breed comes in three forms: smooth-coated, long-haired, and wire-haired.

 

 

Wire-haired dachshund, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

The Samoyed originated among the nomadic Samoyed people in Siberia, to pull sledges and to assist in the herding of reindeer. Today, it is a very popular family dog in the West.

 

 

Samoyed, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

The Rhodesian ridgeback is a large-sized dog breed, which originated as a cross between the ridged hunting dogs of the Khoikhoi people, and European dogs, brought to the Cape Colony of South Africa by the Boer. The name was instigated in 1922 in Bulawayo, South Rhodesia (today Zimbabwe).

 

 

Pup of Rhodesian ridgeback, chewing on a bone, Keetmanshoop, Namibia. Note the ridge along the spine. (Photo copyright © by Kaj Halberg)

 

 

 

The commonest type of stray dogs in Taiwan, generally called Taiwan dogs, or sometimes Takasago dogs, are a result of the indigenous Formosan hunting dogs interbreeding with imported dog types. Taiwan dogs are usually black or brown, or a mixture of the two.

 

 

During Chinese New Year, a red scarf has been tied around the neck of this c. 12-week-old Taiwan pup. A red envelope, on which is written wang-wang, has been fastened to it. The red colour of the envelope, as well as the text, denotes well-wishing. At the same time, wang-wang is an imitation of a dog’s barking. (Photo copyright © by Kaj Halberg)

 

 

Same dog as above, 3 years later. (Photo copyright © by Kaj Halberg)

 

 

 

Lupulella mesomelas Black-backed jackal
In 2017, genetic research concluded that this species and the side-striped jackal (L. adusta) were only distantly related to members of the genus Canis, and, consequently, they were moved to a separate genus.

The black-backed jackal, also known as silver-backed jackal, is quite common in two widely separated areas. The nominate subspecies is distributed in southern Africa, from southern Angola, south-western Zambia, Zimbabwe, and southern Mozambique southwards to the Cape Province of South Africa, whereas subspecies schmidti is found from extreme south-eastern Sudan, Eritrea, and Ethiopia southwards to central Tanzania, westwards to Uganda.

 

 

Resting black-backed jackal, Hwange National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Lycaon pictus African hunting dog
This fascinating animal is described in depth on the page Animals – Mammals: Hunting dogs – nomads of the savanna.

 

 

Hunting dogs are formidable hunters, with powerful jaws and long legs. They are able to run at 60 km/hour. (Photo copyright © by Kaj Halberg)

 

 

 

Vulpes vulpes Red fox
This fox is described in depth on the page Animals – Mammals: Dog family.

 

 

Confiding red fox pup, Copenhagen, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cercopithecidae Old World monkeys
The species below are all described in depth on the page Animals – Mammals: Monkeys and apes.

 

Macaca cyclopis Taiwan macaque

 

 

The fur of Taiwan macaque is pale grey with brownish here and there. These were photographed in the Bagua Shan Mountains, western Taiwan. (Photos copyright © by Kaj Halberg)

 

 

 

Macaca fascicularis Long-tailed macaque

 

 

Long-tailed macaques, Wenara Wana Temple (popularly called ‘Monkey Forest’), Ubud, Bali, Indonesia. (Photos copyright © by Kaj Halberg)

 

 

 

Macaca mulatta Rhesus monkey

 

 

Rhesus monkey, photographed at the Buddhist temple Swayambhunath, Kathmandu, Nepal, where two troops of these monkeys live. (Photo copyright © by Kaj Halberg)

 

 

Female rhesus monkey with a suckling young, Swayambhunath. (Photo copyright © by Kaj Halberg)

 

 

 

Macaca silenus Lion-tailed macaque

 

 

Male lion-tailed macaque, Puthutottam, West Ghats, Tamil Nadu, India. (Photo copyright © by Kaj Halberg)

 

 

 

Papio anubis Olive baboon

 

 

Male olive baboon, Lake Manyara National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Papio cynocephalus Yellow baboon

 

 

Male yellow baboon, Mikumi National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Piliocolobus kirkii Zanzibar red colobus

 

 

Zanzibar red colobus, photographed in Zanzibar. (Photos copyright © by Kaj Halberg)

 

 

 

Semnopithecus schistaceus Nepal langur, pale-armed langur

 

 

The Nepal langur is easily identified by its luxurious, pale grey fur and the large white ruff around its jet-black face. These two were photographed near Lake Dodi Tal, Asi Ganga Valley, Uttarakhand. (Photo copyright © by Kaj Halberg)

 

 

 

Theropithecus gelada Gelada baboon

 

 

In the Simien Mountains, the gelada baboon is fairly common in some areas. This picture shows a female at Gosh Meda. On the chest, males as well as females have a naked red skin patch, giving rise to an alternative name of the species, bleeding-heart monkey. (Photo copyright © by Kaj Halberg)

 

 

 

Trachypithecus auratus Javan lutung

 

 

Javan lutung, for sale at a market in the city of Yogyakarta, Java. (Photo copyright © by Kaj Halberg)

 

 

 

Cervidae Deer
A large number of deer species, including the ones below, are described on the page Animals – Mammals: Deer.

 

Axis axis Spotted deer

 

 

Spotted deer stag, drinking from a waterhole, Sariska National Park, Rajasthan, India. (Photo copyright © by Kaj Halberg)

 

 

 

Cervus canadensis Wapiti

 

 

Resting wapiti stag, Fort Niobrara Wildlife Refuge, Nebraska, United States. (Photo copyright © by Kaj Halberg)

 

 

 

Cervus unicolor Sambar deer

 

 

In Maha Eliya Thenna National Park (Horton Plains), central Sri Lanka, sambar deer have become accustomed to tourists, as this one, which I could approach to within 10 metres. (Photo copyright © by Kaj Halberg)

 

 

 

Delphinidae Oceanic dolphins
This cosmopolitan family contains about 37 species, divided into about 18 genera.

 

Orcinus orca Killer whale, orca
This strikingly patterned dolphin is the largest member of the family, males reaching a length of more than 9 m and weighing up to 10 tons. Females are smaller, up to 7 m long and weighing 3-4 tons. It is the only member of the genus, found in all oceans of the world.

These animals are very intelligent and often perform in ocean parks.

 

 

Killer whale, Ocean Park, Hong Kong. (Photo copyright © by Kaj Halberg)

 

 

 

Echimyidae Spiny rats
A large family with 27 genera and about 100 species, mainly found in South America, with some species in Central America and the Caribbean.

The family name is derived from the Greek ekhinos (‘hedgehog’, ‘sea urchin’) and mys (‘mouse’), alluding to the stiff hairs that many members of the family have on their body, presumably to deter enemies from eating them.

 

Myocastor coypus Nutria, coypu
This animal is native to the southern half of South America, living in wetlands. At an early stage, it was introduced to North America, Europe, and Japan for the fur trade. Over the years, numerous animals have escaped, and the species has become naturalized in many places. It is considered a pest, as it competes with, and sometimes expels, native species, erodes river banks, destroys irrigation channels, and chews up house panels etc.

The generic name is derived from Ancient Greek mys (‘mouse’) and kastor (‘beaver’), alluding to its resemblance to the beavers (Castor).

 

 

Nutria, escaped on Avery Island, Louisiana, United States. (Photo copyright © by Kaj Halberg)

 

 

 

Elephantidae Elephants
Elephants and their sad fate are described on the page Animals – Mammals: Rise and fall of the mighty elephants.

 

Elephas maximus Asian elephant

 

 

Grazing bull Asian elephant, Corbett National Park, Uttarakhand, India. Note that the gland in front of its ear is emitting fluid, indicating that it is in heat, called musht. (Photo copyright © by Kaj Halberg)

 

 

Female Asian elephant in a breeding centre for elephants, near Sauraha, southern Nepal. (Photo copyright © by Kaj Halberg)

 

 

 

Loxodonta africana African elephant

 

 

African elephant, eating grass, Ngorongoro Crater, Tanzania. Note that it has only one tusk. (Photo copyright © by Kaj Halberg)

 

 

After spraying itself, this African elephant is covered in a layer of grey mud as a protection against biting insects, Hwange National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Equidae Horses
The origin of horses, asses, and zebras is described on the page Animals: Animals as servants of Man.

There are 3 species of zebra, living in Africa, 3 species af ass, living in Asia and Africa, and one species of horse, which went extinct in the wild, but has been reintroduced to Mongolia. Herds of feral domestic horses are found many places around the world.

The three zebra species are described on the page Nature: Nature’s patterns.

 

Equus ferus ssp. caballus Domestic horse
In Europe, the true wild horse, or tarpan (Equus ferus ssp. ferus), was hunted to extinction, and the last one died in a Russian zoo in 1909. Another subspecies, Przewalski’s horse (E. ferus ssp. przewalskii), named after Russian geographer and explorer Nikolai Przewalski (1839-1888), only survived as scattered herds on the vast grass steppes in Ukraine, Kazakhstan, and Mongolia. Attempts have been made to recreate the tarpan by crossing Przewalski’s horses with various ancient, primitive types of domestic horses, resulting in tarpan-like horses, such as the Konik horse. Today, herds of these ‘primitive’ horses have been released various places in Europe.

The domestication of wild horses is described on the page Animals: Animals as servants of Man.

 

 

The Konik horse resembles the extinct tarpan, apart from the long mane. A feral population of these horses live in the nature reserve Oostvardersplassen, Flevoland, Holland, where this picture was taken. (Photo copyright © by Kaj Halberg)

 

 

 

As its name implies, the Belgian draft horse originates in Belgium. It is one of the strongest of the heavy horse breeds.

 

 

Portraits of Belgian horses, Bornholm, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Tibetan horses are small and sturdy animals. This one was photographed in the town of Gyantse, Tibet. (Photo copyright © by Kaj Halberg)

 

 

 

A mule is a cross between a jack (male donkey) and a mare (female horse). – Shigatse, Tibet. (Photo copyright © by Kaj Halberg)

 

 

 

Equus quagga Plains zebra

 

 

Resting plains zebra, subspecies chapmani, Hwange National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Previously, plains zebras in Namibia were regarded as a distinct subspecies, antiquorum. However, recent studies have revealed that it is genetically identical to Burchell’s zebra, subspecies burchelli, which was once regarded as extinct. As subspecies burchelli was described prior to antiquorum, the former name takes precedence. Thus, the plains zebras of Namibia are now called Equus quagga ssp. burchelli. Following the extermination of the quagga, ssp. quagga, in the late 1800s, Burchell’s zebra is today the least striped surviving subspecies.

 

 

Burchell’s zebra, mare and foal, Etosha National Park, Namibia. (Photos copyright © by Kaj Halberg)

 

 

 

Erithizontidae New World porcupines
These animals, comprising 19 species in 3 genera, are primarily South American, with a few species extending into Central America, and a single species in North America (below).

 

Erethizon dorsatum North American porcupine
This striking animal, whose body is covered in up to 30,000 quills, is the second-largest rodent in North America, only surpassed by the beaver (Castor canadensis). It grows up to 90 cm long, excluding the long tail, which may be to 30 cm long. It is quite heavy, usually weighing 7-10 kg, sometimes more.

Divided into 7 subspecies, it is distributed in most of subarctic Alaska and Canada, extending its range through western United States to the northernmost parts of Mexico, and in the eastern states it is mainly restricted to the Appalachian Mountains, southwards to Pennsylvania.

The generic name is derived from Ancient Greek erethizein (‘to irritate’), referring to the quills, the specific name from the Latin dorsatus (‘ridged’). Thus, the name can be loosely translated as ‘the animal with the irritating back’.

The word porcupine is derived from the Latin porcus (‘pig’) and spina (‘thorn’). Despite its name, this animal is not closely related to Old World porcupines (Hystrix).

 

 

An adult North American porcupine may have up to 30,000 quills. This one was observed at Bend, Oregon. (Photo copyright © by Kaj Halberg)

 

 

 

Felidae Cats
This worldwide family, only absent from the Australian region, the polar areas, and Madagascar, is divided into two subfamilies, Felinae with about 32 largely smaller species, and Pantherinae with 7 mostly large species.

The snow leopard (Panthera uncia) is described on the page Animals: Animal tracks and traces, whereas the sad fate of the tiger (P. tigris) is related on the page Folly of Man.

 

Acinonyx jubatus Cheetah
The fastest land mammal on Earth, during hunts often running at speeds of up to 64 km/h, being able to accelerate up to 112 km/h on short distances. Because of this ability, the cheetah was tamed as early as the 16th Century B.C. in Egypt, and later also in India, to be used for hunting.

This species mainly inhabits savanna, but is also found in various types of open forest. Four subspecies are currently recognized. The nominate jubatus occurs from Uganda and Kenya southwards through eastern and southern Africa to Namibia and South Africa. It has been exterminated in Zaire, Rwanda, and Burundi. The population is estimated at around 5,000 individuals.

Subspecies soemmeringii is restricted to north-eastern Africa, occurring in South Sudan, Ethiopia, and Eritrea.

With a total population estimated at less than 250 individuals, subspecies hecki is listed as critically endangered by the IUCN. It has a scattered occurrence of tiny populations in southern Algeria, Niger, Burkina Faso, and Benin.

Today, the Asiatic cheetah, subspecies venaticus, is confined to Iran. It is classified as critically endangered by the IUCN, as the total population in 2017 was estimated at fewer than 50 individuals, scattered over the central plateau of Iran. In former times, this subspecies was distributed from the Arabian Peninsula and Turkey eastwards to Central Asia and India.

By 2016, the total cheetah population was estimated at around 7,100 individuals in the wild. Its decline is caused by loss of habitat, poaching for the illegal pet trade, and conflict with humans.

The generic name is derived from the Greek akinitos (‘motionless’) and onyx (‘nail’ or ‘hoof’), thus ‘motionless nails’, referring to the fact that the cheetah, unlike other cats, is unable to retract its claws. The specific name is from the Latin iuba (‘mane’ or ‘crest’) and‎ atus (‘like’), thus ‘having a mane-like crest’, referring to the long mane of cheetah kittens below the age of 3 months. This mane is a means of camouflage, when the kittens are left in dense cover by their mother, when she goes hunting. The name cheetah is derived from the Sanskrit citra, meaning ‘variegated’, ‘spotted’, or ‘speckled’.

 

 

Resting cheetah, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Felis catus Domestic cat
The domestic cat is presented in depth on the page Animals: Animals as servants of Man.

 

 

Large kitten, resting, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Kittens, Funen, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Felis libyca African wildcat
Previously, this small cat was regarded as various subspecies of the European wildcat (F. silvestris), but today most authorities consider it a separate species. Despite its name, it is not only found in Africa. As of today, 3 subspecies are recognized, the nominate in northern Africa, cafra in southern Africa, and ornata in the Arabian Peninsula, the Middle East, north-western India, and Central Asia, eastwards to Mongolia and northern China.

Numerous other subspecies (or even species) have been described from skins that are now regarded as specimens of African wildcat.

 

 

African wildcat, ssp. cafra, resting in savanna grass, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Leopardus pardalis Ocelot
A medium-sized cat, which may grow to almost 1 m long and weighing up to 16 kg. It is distributed from extreme southern Arizona and Texas southwards through western and eastern Mexico and Central America to southern Peru and northern Argentina. Its prime habitat is in the vicinity of water, with dense vegetation cover.

Populations are decreasing in many parts of its range due to habitat destruction, hunting, and traffic accidents.

 

 

Female ocelot in captivity, Tucson Desert Zoo, Arizona. (Photo copyright © by Kaj Halberg)

 

 

 

Panthera leo Lion
Lions live in prides, consisting of females and young, and a single or several males. If there is more than one male, they are brothers or half-brothers. Even if they often don’t participate in a hunt, the stronger males will chase away lionesses and cubs from a prey, if it is not large enough to feed the entire pride.

The lion is unique among cats due to the male’s mane, a large growth of hair around the neck and down the chest, and often a little way down the back. The mane makes a male lion look larger than he actually is, without the disadvantage of a larger weight, which would require more food. A large mane is a signal to other males that here comes a powerful animal that shouldn’t be challenged, even if the challenging male is in fact larger than his opponent, but has a smaller mane. The mane also gives some protection during fights among males, for instance when stray males attempt to take over a pride.

The lion is described in depth on the page Animals – Mammals: Lion – king of the savanna, and an unusual nightly encounter with lions is related on the page Travel episodes – Tanzania 1990: Lions in the camp.

 

 

Male lion, resting among grass, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

This male in Serengeti National Park has a huge mane. (Photo copyright © by Kaj Halberg)

 

 

 

Panthera pardus Leopard
This large cat has a very wide distribution, from Ussuriland in south-eastern Russia southwards through parts of China and Southeast Asia to Indonesia, and thence westwards through the Indian Subcontinent to the Middle East, and most of sub-Saharan Africa. It has adapted to a huge variety of habitats, from tropical jungles to semi-desert and mountains, and even farmland near villages.

More about leopards, including a terrible man-eater from northern India, is found on the page Animals – Mammals: The spotted killer.

 

 

Leopard, Chief’s Island, Okawango, Botswana. (Photo copyright © by Kaj Halberg)

 

 

Leopard, dozing in a tree, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

This leopard is resting after filling its stomach to the bursting point with meat from a white-bearded wildebeest (Connochaetes taurinus ssp. mearnsi), Serengeti National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Giraffidae Giraffe and okapi
This family consists of only two surviving species, the giraffe (below), and the okapi (Okapia johnstoni), which is restricted to eastern Zaire.

 

Giraffa camelopardalis Giraffe
The tall and seemingly ungainly giraffe was once distributed throughout Africa, except in rainforest. However, it has diminished alarmingly, and today there may be less than 100,000 individuals.

According to the International Union for Conservation of Nature (IUCN), only one species of giraffe exists, divided into 9 subspecies, whereas other authorities recognize up to 8 separate species.

The name giraffe was derived from the Arabic zarafah, meaning ‘the one that walks fast’. The specific name is from the Greek words kamelos (‘camel’) and pardos (‘leopard’), and the Latin alis (‘resembling’), alluding to its camel-like shape and leopard-like pattern.

Other pictures, depicting giraffes, are shown on the page Nature: Nature’s patterns.

 

The reticulated giraffe, subspecies reticulata, is distributed in extreme southern Ethiopia, north-eastern Kenya, and south-western Somalia.

 

 

Reticulated giraffes, Buffalo Springs National Park, Kenya. (Photos copyright © by Kaj Halberg)

 

 

 

The Kordofan giraffe, subspecies antiquorum, is found in southern Chad, the Central African Republic, northern Cameroun, and north-eastern Zaire. The pattern somewhat resembles that of the reticulated giraffe, but is much paler. Formerly, it was believed that populations in Cameroun belonged to subspecies peralta, but genetic research has shown that this is incorrect. It is highly endangered, as only about 400 individuals survive in fragmented populations.

 

 

Kordofan giraffe, Waza National Park, Cameroun. (Photo copyright © by Kaj Halberg)

 

 

 

The dark blotches on the Masai giraffe, subspecies tippelskirchi, have a jagged outline. It is distributed in Tanzania and extreme southern Kenya.

 

 

Masai giraffe, Mikumi National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

Masai giraffe, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

The giraffes in Arusha National Park, northern Tanzania, are probably Masai giraffes, although some authorities regard them as hybrids between reticulated giraffe and Masai giraffe, called Galana giraffes. (Photo copyright © by Kaj Halberg)

 

 

 

The Angolan giraffe, subspecies angolensis, is found in south-western Zambia, central Botswana, Zimbabwe, and northern Namibia.

 

 

Angolan giraffe, feeding on acacia leaves, Chief’s Island, Okawango, Botswana. (Photo copyright © by Kaj Halberg)

 

 

Young Angolan giraffes, Chief’s Island. (Photo copyright © by Kaj Halberg)

 

 

 

Hippopotamidae Hippos
A small family of only two surviving species. The hippo (below) and its smaller cousin, the pygmy hippo (Choeropsis liberiensis), are both described on the page Animals – Mammals: Hippo – the river horse that lives on both sides.

 

Hippopotamus amphibius Hippo

 

 

Hippos can crush a canoe with a single bite from their enormous jaws. – Ngorongoro Crater (top), and Banagi River, Serengeti National Park, both in Tanzania. (Photos copyright © by Kaj Halberg)

 

 

 

Hominidae Man and apes
Previously, apes (orangutans, gorillas, and chimpanzees) were placed in the family Pongidae.

 

Pongo Orangutans
Formerly, orangutans were regarded as a single species, Pongo pygmaeus, which was confined to rainforests of Sumatra and Borneo. Lately, however, it has been split into three separate species. The Bornean orangutan (P. pygmaeus), comprising 3 subspecies, is still widespread in Borneo, whereas the Sumatran orangutan (P. abelii) is restricted to northern Sumatra, primarily the Aceh Province. The Tapanuli orangutan (P. tapanuliensis), which lives in an area of just 1,500 km2 in Batang Toru, south of Lake Toba, Sumatra, was described as a distinct species as late as 2017. They all live in lowland rainforest, rarely found above 500 m altitude, the Sumatran orangutan occasionally up to c. 1,500 m.

All three species are seriously endangered due to habitat destruction and fragmentation, and illegal poaching for zoos. It has been estimated that the population of the Bornean orangutan in 1973 was about 300,000 individuals, and it is assumed that this number will decline to c. 50,000 by 2025. The Sumatran orangutan has an estimated population of fewer than 14,000, and it is predicted that this number will decline by 80% by 2060. The population of Tapanuli orangutan is fewer than 800, and this number is still decreasing.

 

Pongo pygmaeus Bornean orangutan
In 1985, I visited the Sepilok Orangutan Rehabilitation Centre in Sabah, Borneo. This centre is home to a number of orphaned orangutans, which were confiscated from poachers, who shot their mothers to get hold of the young. This centre, where the young ones are trained to a life in the wild, is presented in depth on the page Travel episodes – Borneo 1985: Visiting orangutans.

 

 

An adult male, living in a semi-wild state near the centre. (Photo copyright © by Kaj Halberg)

 

 

Lacking a mother, the orphaned young often become much attached to one another. (Photo copyright © by Kaj Halberg)

 

 

 

Hyaenidae Hyaenas
About 5 million years ago, this family was much larger, containing at least 21 genera with more than 50 species, widely distributed in Eurasia, Africa, and North America. Today, only 4 species remain: spotted hyaena (below), striped hyaena (Hyaena hyaena), brown hyaena (Parahyaena brunnea), and aardwolf (Proteles cristatus). They are restricted to Africa, with the exception of the striped hyaena, which is also found in the Middle East and India.

 

Crocuta crocuta Spotted hyaena
This powerful carnivore is found in most of sub-Saharan Africa, with the exception of deserts, rainforests, and alpine areas on mountain tops. It once ranged all over Europe and northern Asia, from Spain and France eastwards to eastern Siberia. It is still not clear why it went extinct in Siberia, but its disappearance from Europe is linked to the decline in grasslands – its favoured habitat – about 12,500 years ago.

The spotted hyaena has a very complex social behaviour, with respect to group-size, hierarchical structure, and frequency of social interaction among both kin and unrelated group-mates. However, their social system is openly competitive rather than cooperative, with access to kills, mating opportunities, and the time of dispersal for males, all depending on the ability to dominate other clan-members. (Source: Holekamp, Sakai & Lundrigan, 2007. Social intelligence in the spotted hyena (Crocuta crocuta). Philosophical Transactions of the Royal Society, London, 362, pp. 523-538)

 

 

Resting spotted hyaena, Serengeti National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Leporidae Hares and rabbits
This family contains more than 60 species, of which about 32 belong to the genus Lepus (true hares). Members are found on all continents, except Antarctica, although they have been introduced to Australia. The family name means ‘those that resemble lepus’, lepus being the classical Latin word for hare.

 

Lepus europaeus European hare
This species is native to the major part of Europe and the Middle East, and thence eastwards across the Asian steppes to Mongolia. It has also been introduced elsewhere, including Britain, Australia, New Zealand, and southern South America.

 

 

European hare, resting in a littoral meadow, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Oryctolagus cuniculus European rabbit
Originally, this species was native to south-western Europe and north-western Africa, but has been introduced to many other areas as a hunting object. It was first brought to Britain by the Romans, following their invasion in A.D. 43, and today it is extremely common – an estimate in 2004 suggested about 40 million.

In Australia, 24 European rabbits were introduced in Victoria in 1859. The vast farmlands were an ideal habitat for the rabbits, and the mild winters allowed them to breed year-round, so they quickly spread over most of the country. Australia’s equivalent to the rabbit, the greater bilby (Macrotis lagotis), could not compete with the fast-breeding rabbits, and today it is an endangered species.

Other places of introduction include New Zealand, certain Hawaiian islands, and islands off the coast of South Africa. There are also many instances of pet rabbits, which have escaped, forming feral populations.

In countless cases, this species has done severe damage to the environment, partly through overgrazing, partly through its system of underground tunnels, and partly through competition with local wildlife. It is regarded as an invasive species in most countries.

In 1972, English author Richard Adams (1920-2016) wrote a novel, Watership Down, describing the life of a group of wild rabbits in southern England. Although the rabbits have been anthropomorphized, the story gives a fair account of the life of these animals.

 

 

The European rabbit was introduced to Britain almost 2,000 years ago, and today it is extremely common. This one is sitting outside its den in Spey Valley, Scotland. (Photo copyright © by Kaj Halberg)

 

 

 

Lorisidae Lorises, pottos, and angwantibos
This family of small, nocturnal primates contain 5 genera with about 16 species, living in South and Southeast Asia, and in Africa south of the Sahara.

 

Nycticebus menagensis Philippine slow loris
This species, which was previously regarded as a subspecies of the Sunda slow loris (N. cougang), is native to northern and eastern Borneo, and the Sulu Archipelago in the Philippines. It is nocturnal and arboreal, living in evergreen forests.

 

 

This Philippine slow loris had been injured and was now recovering in Sepilok Orangutan Rehabilitation Centre, Sabah, Borneo. (Photo copyright © by Kaj Halberg)

 

 

 

Otariidae Eared seals
As their name implies, these seals have a small ear flap, which distinguishes them from the true seals, family Phocidae, and the walrus (Odobenus rosmarus). Eared seals include sea lions and fur seals, altogether 7 genera with 15 species, occurring throughout the Pacific Ocean and the southern parts of the Indian and Atlantic Oceans. They are absent from the north Atlantic.

 

Arctocephalus pusillus Brown fur seal
There are two widely separated populations of this seal, also known as Afro-Australian fur seal: the South African, or Cape, fur seal, subspecies pusillus, and the Australian fur seal, subspecies doriferus.

The Cape fur seal ranges along the southern coasts of Africa, from Ilha dos Tigres in southern Angola, along the Namibian coast to Algoa Bay in South Africa, whereas the Australian subspecies lives in south-eastern Australian waters, along the coasts of Tasmania, New South Wales, Victoria, and South Australia, with the largest concentration in the Bass Strait.

The preferred breeding habitats of these seals are rocky islands, or pebble or boulder beaches. The population of the Cape fur seal is approximately 2 million, whereas that of the Australian fur seal is around 120,000. (Source: iucnredlist.org/details/2060/0)

 

 

Bull Cape fur seal, surrounded by females and pups, Cape Cross, Namibia. (Photo copyright © by Kaj Halberg)

 

 

Sleeping female Cape fur seal, Cape Cross, Namibia. (Photo copyright © by Kaj Halberg)

 

 

 

Eumetopias jubatus Northern sea lion
This animal, also known as Steller’s sea lion, lives in the northern Pacific, from the Kuril Islands in Russia to the Gulf of Alaska in the north, and thence southwards to central California. It is the largest of the eared seals (Otariidae) and the sole member of the genus Eumetopias.

The name Steller’s sea lion commemorates German naturalist, physician, and explorer Georg Wilhelm Steller (1709-1746), who described the species in 1741. He participated in the Second Russian Kamchatka Expedition, led by Danish explorer Vitus Bering (1681-1741).

 

 

Female northern sea lion, Oregon, United States. (Photo copyright © by Kaj Halberg)

 

 

 

Procaviidae Hyraxes
Hyraxes constitute two genera, Procavia and Dendrohyrax, in Procaviidae, the only living family within the order Hyracoidea. These animals resemble large guinea pigs, but their nearest living relatives are in fact elephants.

 

Dendrohyrax arboreus Southern tree hyrax
The 3 species of tree hyraxes, genus Dendrohyrax, are distributed in sub-Saharan Africa. The southern tree hyrax is found from eastern Zaire, southern Uganda, and southern Kenya southwards to eastern Angola, Zambia, and northern Mozambique, with two isolated populations in southern Mozambique and south-eastern South Africa. This animal lives in various types of forest, and also in savanna and rocky areas, provided there are trees. It may be encountered from the lowlands up to an elevation of 4,500 m.

 

 

Southern tree hyrax, Ngorongoro Crater, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Procavia capensis Rock hyrax
Divided into at least 5 subspecies, this animal is distributed in most of northern Africa, from southern Algeria, southern Libya, and the Nile Valley southwards to Zaire and northern Tanzania, in the Arabian Peninsula, and in southern Africa. In South Africa, it is known as dassie. It lives in rocky areas, from sea level up to elevations around 4,200 m. Where it is regularly fed, it can become very tame.

 

 

Black-necked rock hyrax, subspecies johnstoni, scratching, Gorges Valley, altitude about 4,000 m, Mount Kenya. (Photo copyright © by Kaj Halberg)

 

 

 

Rhinocerotidae Rhinos
A small family of 5 species in 4 genera, distributed in eastern and southern Africa, northern India, southern Nepal, Indochina, and Indonesia.

Rhino parts have been used as ingredients in traditional Asian medicine for at least 2000 years. Virtually every part of the animal is used: the horn for reducing fever and spasms; the skin for skin diseases; the penis as an aphrodisiac; the bones to treat bone disorders; the blood “as a tonic for women who are suffering from menstrual problems.”

In China, powdered horn is regarded as an aphrodisiac. However, chemical analyses have not revealed any active ingredients to suggest that the remedy could be effective in this respect. (Source: J. Still 2003. Use of animal products in traditional Chinese medicine. In: Complementary Therapies in Medicine)

In fact, western medical experts tend to discount all claims of any curative power in rhino horn. It is well known that aspirin contains similar properties and produces many of the same results as rhino prescriptions in patients.

Formerly, rhino horn was used for adorning dagger sheaths in Yemen – a practice which may still take place.

All five species of rhino are critically endangered due to widespread poaching, the Asian species also due to habitat loss.

 

Ceratotherium simum White rhinoceros, square-lipped rhinoceros
This is the largest living species of rhino, growing to 4 m long and weighing up to 2.3 tonnes. Females live in small herds, as opposed to other rhinos, which are largely solitary. There are two subspecies, the southern nominate race, which counts about 20,000 individuals, and the northern, subspecies cottoni, which has gone extinct in the wild due to poaching. Only two animals survive in captivity.

The generic name is derived from Ancient Greek keras (‘horn’) and therion (‘beast’). The specific name is derived from the Greek simos (‘snub-nosed’), alluding to the square mouth of this species, an adaptation for grazing.

It has often been claimed that the most commonly used name, white rhino, is a mistranslation of the Dutch word wijd to the English word white. Wijd means ‘wide’ in English, and it was supposed to refer to the width of the rhinoceros’s mouth. However, this is not the case. In fact, the name white rhino can be traced back to a letter in Dutch, written by the Boer Petrus Borcherds to his father in 1802. In this letter, he mentions two rhinos, both killed in 1801, a male of the ‘black variety’, and a female ‘white’ rhino. Concerning the female, Borcherds stated (still in Dutch): “She was of the type known to us as the white rhinoceros. (…) I expected this animal to be entirely white, according to its name, but found that she was a paler ash-grey than the black male.” (Source: Jim Feely 2007. Black rhino, white rhino: what’s in a name? Pachyderm 43, pp. 111-115)

However, both species are in reality grey, the ‘black’ rhino somewhat darker than the ‘white’ rhino.

 

 

White rhino, Matobo National Park, Zimbabwe, wearing a radio collar for easy tracking. Its horn was removed to deter poachers from killing it, but has started growing out again. The number of rhinos in Zimbabwe has plummeted in later years due to poaching, and the present population is only 7-800 individuals. (Photo copyright © by Kaj Halberg)

 

 

 

Diceros bicornis Black rhinoceros, hook-lipped rhinoceros
In former days, this rhino was abundant in sub-Saharan Africa, divided into 7 or 8 subspecies. However, due to poaching it has largely disappeared, today surviving in small populations in reserves in Kenya, Tanzania, and southern African countries.

Both scientific names mean ‘two-horned’, the generic name derived from the Greek dyo (‘two’) and keras (‘horn’), the specific name from the latin bis (‘twice’) and cornu (‘horned’). The common name is explained above, see white rhino.

 

 

Resting black rhinos, Ngorongoro Crater. (Photos copyright © by Kaj Halberg)

 

 

 

Rhinoceros unicornis Indian rhinoceros, greater one-horned rhinoceros
Formerly, this animal was widespread and common in grasslands of northern India and southern Nepal, but today only about 3,500 survive in small pockets, with about 70% of the entire population in Kaziranga National Park, Assam.

The generic name is derived from Ancient Greek rhinos (‘nose’) and keras (‘horn’), the specific name from the Latin unus (‘one’) and cornu (‘horned’).

 

 

Indian rhino, Kaziranga National Park, Assam, India. (Photo copyright © by Kaj Halberg)

 

 

This Indian rhino is enjoying a mudbath, near Sauraha, southern Nepal. (Photo copyright © by Kaj Halberg)

 

 

 

Sciuridae Squirrels
A large family of rodents, comprising about 60 genera and c. 300 species worldwide. These animals are found on all continents, except Antarctica. In Australia, however, they have been introduced by humans.

Numerous members of the family are described on the page Animals – Mammals: Squirrels.

 

Callosciurus
This genus contains about 15 species, found mainly in Southeast Asia, with a few species occurring in Nepal, northeastern India, Bangladesh, southern China, and Taiwan. Most species live in tropical rain forests, but some have adapted to a life in city parks and gardens. Their food consists of nuts, fruits, and seeds, and also insects and bird eggs.

The first part of the generic name is derived from the Greek kallos (‘beautiful’) – although many of the species are of a rather plain brown colour.

 

Callosciurus erythraeus Red-bellied squirrel, Pallas’s squirrel
Despite its name, this species does not always have a red belly. It is widespread, found from eastern Nepal, Bhutan, and north-eastern India eastwards to Southeast Asia, southern China, and Taiwan. More than 30 subspecies have been described, although not all are recognized by many authorities.

 

 

Subspecies taiwanensis of the red-bellied squirrel is common and widespread at lower elevations in Taiwan. This one was feeding on bird food near a temple, dedicated to Confucius, in Tainan, southern Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Marmota Marmots
Marmots are large, ground-living squirrels, comprising 15 species, which occur in Europe, Asia, and North America. They are the largest and fattest of squirrels, growing up to 60 cm long and weighing up to 7 kg.

Marmots live in burrows up to 2 m underground, digging tunnels up to 10 m long. They often sit on their haunches outside their burrow, and their shrill warning whistles can be heard far away. Marmots are almost exclusively vegetarians. They do not make food deposits, but in autumn they have become extremely fat, and hibernate throughout the winter.

 

Marmota marmota Alpine marmot
This is a common animal, living in alpine areas at elevations between 800 and 3,200 m, in southern France, the Vosges, the Black Forest, the Alps, the northern Apennines, the Carpathians in Romania, and the Tatras in Poland. In 1948, it was reintroduced with success in the Pyrenees, where it had disappeared at end of the last Ice Age.

 

 

Alpine marmot outside its den, Gran Paradiso National Park, Italy. (Photo copyright © by Kaj Halberg)

 

 

 

Ratufa Oriental giant squirrels
A genus with 4 species of large squirrels, distributed from India and Sri Lanka eastwards to Southeast Asia and southern China, and thence southwards to Indonesia.

 

Ratufa indica Indian giant squirrel, Malabar giant squirrel
This species is endemic to India, mainly found in forests and woodlands in hilly areas. It is strictly arboreal, and most of its food consists of leaves, fruits, and seeds.

 

 

Indian giant squirrel, feeding on pulp of a species of breadfruit, Artocarpus hirsutus, Periyar National Park, Kerala, South India. (Photo copyright © by Kaj Halberg)

 

 

 

Suidae Pigs
Members of this family, counting 6 genera with 18 or 19 species, are native to Europe, Asia, and Africa. New World pigs, called peccaries or javelinas, belong to a different family, Tayassuidae.

 

Sus (scrofa) domesticus Domestic pig
Some authorities recognize the domestic pig as a separate species, others regard it as a subspecies of the wild boar (Sus scrofa). The domestic pig is described in depth on the page Animals: Animals as servants of Man.

 

 

Free-running pigs, Fur, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Free-running pigs, wallowing in mud, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Phacochoerus africanus Common warthog
Previously, this animal was thought to be the only member of the subfamily Phacochoerinae, under the name P. aethiopicus, but recently it has been split into two species, the desert warthog, named P. aethiopicus, which lives in arid areas of northern Kenya, Somalia, and eastern Ethiopia, and the widespread common warthog, named P. africanus, which lives in grassland and woodland in most of sub-Saharan Africa, only avoiding deserts and rainforest.

From a distance, this animal appears largely naked, seemingly only with a crest along the back, and tufts of hair on the cheeks and tail. At close quarters, however, you notice a cover of short, bristly hairs on the body. The name warthog refers to the facial wattles, larger in the males, which also have prominent tusks that may sometimes reach a length of up to 60 cm, much smaller in the females.

 

 

Male common warthog with huge tusks and prominent wattles, Hwange National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

Female common warthog with very small tusks and almost no wattles, Queen Elizabeth National Park, Uganda. (Photo copyright © by Kaj Halberg)

 

 

 

Ursidae Bears
There are altogether 8 species of bears, widely distributed in Eurasia and North America, and a single species, the spectacled bear (Tremarctos ornatus), in the Andes of South America.

Due to illegal hunting, and to feed the insatiable Asian markets for traditional medicine and bear paw soup, bears have disappeared, or become very rare, in many areas, including Europe, Southeast Asia, Korea, China, and Taiwan. Today, five species are endangered.

 

Ursus thibetanus Asian black bear, moon bear
This bear is distributed at scattered locations, from south-eastern Iran eastwards across the Himalaya and Indochina to China, Taiwan, Korea, Japan, and south-eastern Siberia (Ussuriland). Previously, it was found in a much larger area, but has declined drastically.

Many Asian black bears are kept in captivity to supply the markets for traditional medicine and bear paw soup. In South Korea, for instance, only around ten individuals live in the wild, whereas about 1,600 are kept in captivity, often under horrible conditions. These captive bears are often killed in the most cruel and horrendous ways, and that this practice is illegal does not seem to deter consumers.

 

 

Asian black bear, photographed in Chengdu Zoo, Sichuan Province, China. (Photo copyright © by Kaj Halberg)

 

 

 

Birds

 

Accipitridae Eagles, hawks, and allies

 

Gyps bengalensis Indian white-rumped vulture
This bird was once abundant in the Indian Subcontinent, but during the last 30 years or so, most populations of this species, and other vultures, have diminished alarmingly due to poisoning from diclofenac, a veterinary drug widely used to treat diseases in livestock. Research has shown that when these vultures feed on cattle carcasses, diclofenac causes kidney failure.

Today, it is critically endangered, and maybe as few as 6,000 individuals exist in the wild.

 

 

This Indian white-rumped vulture is being teased by a house crow (Corvus splendens), which is pulling its feathers, Jaisalmer, Rajasthan. (Photo copyright © by Kaj Halberg)

 

 

 

Haliaeetus vocifer African fish-eagle
This iconic bird is widely distributed in sub-Saharan Africa, always near water. It has been declining in later years, probably due to widespread usage of pesticides.

It was described by French naturalist François Levaillant (1753-1824), who named it vocifer (’the one who has a penetrating voice’) – a most suitable name for this eagle, whose scream is often resounding over the African landscape. It is the national bird of no less than three countries: Zambia, Zimbabwe, and South Sudan.

 

 

This African fish-eagle, observed at the Rufiji River, Tanzania, was indeed confiding. Our rubber dinghy bumped into the tree, in which it was resting, but it merely glanced down at us and didn’t take off. (Photo copyright © by Kaj Halberg)

 

 

 

Alaudidae Larks
A family with 21 genera and about 100 species, distributed in Africa and Eurasia, with a single species reaching the Americas, and Australia, respectively.

 

Alauda arvensis Skylark
This bird breeds in a vast area, from the entire Europe eastwards to Kamchatka, Korea, and Japan, southwards to northern Africa, Turkey, northern Iran, Mongolia, and northern China. Northern populations are migratory, wintering as far south as northern Africa, Arabia, Pakistan, and southern China.

 

 

This exhausted skylark was found in Nature Reserve Tipperne, Ringkøbing Fjord; Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Alcedinidae Kingfishers
Kingfishers, comprising about 114 species of small to medium-sized, often brilliantly coloured birds, are characterized by having a large head, a long, sharp, pointed bill, and very short legs. As their name implies, most of these birds eat fish, although many species live away from water, eating mainly small invertebrates.

These birds are divided into 3 subfamilies: river kingfishers (Alcedininae), tree kingfishers (Halcyoninae), and water kingfishers (Cerylinae).

 

Halcyon
A genus of 11 medium-sized kingfishers, distributed in warmer areas of Africa and Asia.

The generic name is associated with a bird of Greek legend, called Halcyon, generally thought to be the common kingfisher (Alcedo atthis). The Ancients believed that this bird made a floating nest in the Aegean Sea and had the power to calm the waves while brooding her eggs. Two weeks of calm weather were to be expected when the Halcyon was nesting, which took place around winter solstice. These Halcyon days were generally regarded as beginning on the 14th or 15th of December.

This belief in the bird’s power to calm the sea originated in a myth recorded by Roman poet Publius Ovidius Naso (43 B.C. – c. 17 A.D.), known as Ovid. The story goes that Aeolus, ruler of the winds, had a daughter named Alcyone, who was married to Ceyx, the king of Thessaly. Ceyx drowned at sea, and in her grief, Alcyone threw herself into the waves. However, instead of drowning, she was transformed into a bird and carried to her husband by the wind. (Source: phrases.org.uk)

 

Halcyon albiventris Brown-hooded kingfisher
This bird is very widely distributed in Sub-Saharan Africa, from Congo eastwards to southern Somalia and thence southwards to Namibia and South Africa. It lives in woodland, shrubberies, grasslands with trees, parks, and gardens, and may also be observed in cultivated areas. It occurs from the lowland up to an elevation of about 1,800 m.

 

 

Brown-hooded kingfisher, Msumbugwe Forest, northern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Halcyon leucocephala Grey-headed kingfisher
A bird of various types of forest, especially riverine woodland. Nests have been found in riverbanks. It is quite common south of the Sahara, from Mauritania and Senegal eastwards to Ethiopia and Somalia, and thence southwards to South Africa, although absent from the southernmost part. It is also found on the Cape Verde Islands and in southern Arabia.

The specific name is derived from Ancient Greek leukos (‘white’) and kephale (‘head’), although the head is grey rather than white.

 

 

Grey-headed kingfisher, Dar es Salaam, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Ispidina
This genus contains only 2 African species, closely related to members of the genus Corythornis.

The generic name is from the Latin hispida (‘kingfisher’).

 

Ispidina picta African pygmy kingfisher
This tiny kingfisher is widespread in forested areas south of the Sahara, being absent from the Somali Desert and large parts of south-western Africa.

The specific name is Latin, meaning ‘painted’, alluding to the colourful plumage of this bird.

 

 

African pygmy kingfisher, Rondo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Anatidae Ducks, geese, and swans
At present, this large family contains 43 genera with about 146 species, distributed almost worldwide.

 

Anas
Following genetic studies, this formerly very large genus has been divided into 7 genera, with 31 species remaining in Anas. The generic name is the classical Latin word for duck.

 

Anas platyrhynchos Mallard
A very widely distributed species, found across subarctic, temperate, and subtropical areas of North America and Eurasia, southwards to Mexico, Morocco, Egypt, Pakistan, and China. It has also been introduced to many other places as a hunting object, including South America, New Zealand, Australia, and South Africa.

The drake is a gorgeous bird in breeding plumage, with grey sides, purple breast, and glossy-green head, with a blue shine from certain angles. A picture of the female, which is a uniform speckled brown, is shown on the page Animals – Animals as servants of Man: Poultry.

 

 

Mallard drake, feeding in a moat at Christiania, Copenhagen. A water plant is draped over its back. (Photo copyright © by Kaj Halberg)

 

 

 

Anser Grey geese
A genus of 11 species, distributed in arctic and temperate areas of the Northern Hemisphere. The generic name is the classical Latin word for geese.

 

Anser brachyrhynchus Pink-footed goose
This goose has two separate populations, one breeding in eastern Greenland and Iceland, the other one in Svalbard. The western population spends the winter in the United Kingdom and Ireland, whereas the Svalbard population winters in the Netherlands, Germany, and Denmark.

Over the last 50 years, the population has greatly increased due to protection from shooting on the wintering grounds. Numbers in Ireland and England have risen from about 30,000 in 1950 to about 300,000 today.

The pink-footed goose is closely related to the bean goose (A. fabalis) and was formerly treated as a subspecies of that species. The specific name is derived from Ancient Greek brakhys (‘short’) and rhynkhos (‘snout’, ‘bill’), alluding to the rather short bill of the bird.

 

 

Gosling of pink-footed goose, Mjoidalur, northern Iceland. (Photo copyright © by Kaj Halberg)

 

 

 

Aythya
A genus of 12 species, chiefly found in the Northern Hemisphere, with one species in New Guinea, Australia, New Zealand, and on some Pacific islands, another one in New Zealand, and one in Madagascar. Two species are critically endangered, the East Asian Baer’s pochard (A. baeri) and Madagascar pochard (A. innotata).

 

Aythya fuligula Tufted duck
As a breeding bird, this species has an extremely large distribution, found from the entire Europe eastwards to the Pacific Ocean, southwards to the Mediterranean, Kazakhstan, Mongolia, and northern China. Northern populations are migratory, wintering as far south as northern and eastern Africa, the Indian Subcontinent, Indochina, China, Japan, and the Philippines.

 

 

Male tufted duck, encountered in a city park, Århus, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cygnus Swans
A genus of 6 long-necked birds, distributed on all continents except Antarctica. Genetic research indicates that the coscoroba swan (Coscoroba coscoroba) of South America is not a true swan, but more closely related to geese or shelducks.

The generic name is a Latinized form of the Classical Greek word for swans, kyknos.

 

Cygnus cygnus Whooper swan
A widespread and common bird, breeding from Iceland eastwards across Scandinavia and northern Asia to Kamchatka, southwards to southern Sweden, Kazakhstan, Mongolia, and extreme northern China. It is an irregular and scarce breeding bird in Ireland, Scotland, Germany, and Denmark. It is partly resident in Iceland and Norway, but the vast majority of the population is migratory, spending the winter in western and southern Europe, around the Black, Caspian, and Aral Seas, in eastern China, and in Korea and Japan.

As opposed to the mute swan (C. olor), this swan is very vocal, reflected in its English name, which, incidentally, is pronounced ‘hooper swan’, although the word whooper alone is pronounced ‘wooper’. In his book Ornithologiae libri tres (‘Ornithology, Book Three’), from 1676, English naturalist Francis Willughby (1635-1672) refers to the whooper swan as “the Elk, Hooper, or wild Swan”.

The mute swan is described on the page Animals: Urban animal life.

 

 

Whooper swan, grazing on land, Lake Hornborga, Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Mareca Wigeons and allies
A small genus of 5 species, comprising 3 species of wigeon, the gadwall (M. strepera), and the falcated teal (M. falcata). Previously, they were all included in the genus Anas.

The generic name is from a Brazilian-Portuguese word, marréco, meaning ‘small duck’.

 

Mareca penelope Eurasian wigeon
In breeding plumage, the male Eurasian wigeon is a very pretty bird, whereas the female is a uniform pale brown, with darker wings and whitish belly.

It breeds in Temperate Eurasia, from Iceland and Scotland eastwards to the Pacific, whereas the wintering quarters encompass the major part of Europe and the Middle East, northern and eastern Africa, the Indian Subcontinent, Indochina, southern China, Taiwan, and the northern Philippines.

In North America, it is a regular winter visitor to the Pacific lowlands, from Canada southwards to California. Small numbers are also seen elsewhere in the continent.

This species was named Anas penelope by Swedish naturalist Carl Linnaeus (1707-1778) in 1758. He may have referred to Penelope, who, in Homer’s Odyssey, was the wife of Odysseus. She was known for her fidelity to him, while he was away on his journeys. However, as most ducks are certainly not known for their fidelity, this connection is hard to discover.

Another possible explanation may be the Proto-Greek word penelops, mentioned as ‘some kind of bird’ by Greek grammarian Hesychius of Alexandria, who, probably in the 5th or 6th century A.D., compiled a lexicon of unusual and obscure Greek words. (Source: R. Beekes 2009. Etymological dictionary of Greek. Brill)

 

 

Male wigeon, Anadyr, Chukotka, eastern Siberia. (Photo copyright © by Kaj Halberg)

 

 

 

Anhingidae Darters
Darters, also called snakebirds due to their long, thin, flexible neck, are large water birds, comprising two or four species in the genus Anhinga, the sole genus of the family. The American darter (A. anhinga), often called anhinga, lives in the New World.

One or three species are found in the Old World. If only one species is acknowledged, it is called A. melanogaster. Most authorities, however, recognize three full species: Oriental (below), African (A. rufa), and Australasian (A. novaehollandiae).

The generic name means ‘little head’ in the Tupi language of Brazil, referring to an evil spirit of the forests, the devil bird. (Source: G. Marcgrave 1648. Historiae rerum naturalium Brasiliae. Liber V)

 

Anhinga melanogaster Oriental darter
This species breeds in a vast area, found in the Indian Subcontinent, and from Indochina and the Philippines southwards to Indonesia.

 

 

Oriental darter, drying its wings, Keoladeo National Park, Rajasthan, India. (Photo copyright © by Kaj Halberg)

 

 

 

Aramidae

 

Aramus guarauna Limpkin
This bird, the only member of the family, is related to rails and cranes. It lives in marshy areas, eating mainly molluscs, especially apple snails (Pomacea). It is distributed from Florida, the Caribbean, and southern Mexico southwards to northern Argentina.

The generic name refers to a kind of heron, mentioned by Hesychius of Alexandria, a Greek grammarian who, probably in the 5th or 6th Century A.D., compiled a work, titled Alphabetical Collection of All Words, a list of unusual and obscure Greek words. The specific name is the name of a kind of marshbird in the Tupi language, a people who once lived in Brazil. The popular name refers to the fact that the bird seems to limp when it walks.

 

 

Limpkin, Corkscrew Swamp Sanctuary, Florida. (Photo copyright © by Kaj Halberg)

 

 

 

Ardeidae Herons, egrets, and bitterns
Herons comprise about 64 species of long-legged and long-beaked, fish-eating water birds, distributed almost worldwide. Some species are called egrets, mainly birds with ornate plumes during the breeding season, whereas birds of the genera Botaurus, Ixobrychus, and Zebrilus are called bitterns.

 

Ardea
A genus with about 13 species of mainly large herons, distributed almost worldwide. The generic name is the classical Latin word for heron.

 

Ardea cinerea Grey heron
The grey heron is distributed in the major part of Eurasia and Africa, and on Madagascar. Other pictures, depicting this bird, are shown on the pages Fishing, and Animals: Urban animal life.

 

 

Grey heron, feeding in a moat at Christiania, Copenhagen. (Photo copyright © by Kaj Halberg)

 

 

 

Gorsachius melanolophus Malayan night-heron
As a breeding bird, this species has a scattered occurrence, found in north-eastern and south-western India, in southern China and Indochina, and in Taiwan and the Philippines. The majority of the populations are resident, but north Indian and some Chinese birds are migratory, spending the winter in Malaysia and Indonesia. It also occurs in Sri Lanka in winter.

The main habitats of this bird are forests and marshy areas, but in Taiwan, where it is common, it is often found in city parks, where it can become remarkably confiding.

The generic name is derived from the name Ardea goisagi, which Dutch zoologist and museum director Coenraad Jacob Temminck (1778-1858) applied in 1836 to the Japanese night-heron (today called Gorsachius goisagi). In fact, Goi-sagi is the Japanese name for the black-crowned night-heron (below), derived from goi (‘fifth rank’) and sagi (‘heron’). In Japanese classical folklore, the all-powerful Emperor Daigo (reigned 897-930 A.D.) ordered a vassal to capture a black-crowned night-heron. Upon hearing the imperial command, the heron submitted itself to capture. The emperor was pleased that the heron had confirmed his omnipotence over nature as well as man, whereupon he granted it the title ‘king of the herons’ and the position of fifth rank in his court, and released it unharmed. (Source: J.A. Jobling, 2010. The Helm Dictionary of Scientific Bird Names, Christopher Helm, London)

The specific name is derived from Ancient Greek melas (‘black’) and lophos (‘crest’).

 

 

Malayan night-heron, Xitun Botanical Garden, Taichung, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Nycticorax nycticorax Black-crowned night-heron
This smallish heron is found in most warmer parts of the world, also in Taiwan, where it is a common breeding bird in the lowlands.

It figures in Japanese classical folklore (see Malayan night-heron above).

The generic (and specific) name is derived from the Greek nyktos (‘night’) and korax (‘raven’), referring to the mainly nocturnal feeding habits of these birds, and to their hoarse, raven-like call.

 

 

This black-crowned night-heron was ill, sitting on a jogging path near the Han River, Taichung, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Chloropseidae Leafbirds
This family consists of a single genus, Chloropsis, with 13 species of attractive bright green or turquoise passerines, distributed from the Indian Subcontinent eastwards to southern China, and thence southwards through Indochina and the Philippines to Indonesia.

The family and generic names are derived from Ancient Greek khloros (‘green’) and opsis (‘appearance’).

 

Chloropsis jerdoni Jerdon’s leafbird
This colourful bird is widely distributed in the Indian Subcontinent. The male differs from the female by having a black throat and a blue moustachial stripe.

Previously, Jerdon’s leafbird was regarded as a subspecies of the more widespread blue-winged leafbird (then called C. cochinchinensis). However, it differs in morphology, lacking the blue flight feathers of that species. Following recent genetic research, C. cochinchinensis has been split into 3 separate species.

The specific name commemorates British physician and naturalist Thomas Caverhill Jerdon (1811-1872), who described many birds species in India, several of which are named after him.

 

 

This male Jerdon’s leafbird was caught in a mistnet to be ringed, near Mysore, Karnataka, south India. (Photo copyright © by Kaj Halberg)

 

 

 

Ciconiidae Storks
These large birds, comprising 6 genera with 19 species, are found in most parts of the world, being absent from the polar regions and the major parts of North America and Australia.

The family name is derived from ciconia, the Classical Latin word for storks.

 

Leptoptilos
A genus of 3 large and ungainly birds, found in sub-Saharan Africa and tropical Asia. The generic name is derived from Ancient Greek leptos (‘thin’ or ‘slender’) and ptilon (‘feather’), alluding to the scarcity of feathers on the head of these birds.

 

Leptoptilos crumenifer Marabou stork
This huge bird, up to 1.5 m tall and weighing up to 8 kg, is not exactly a beauty, with a naked, red and black head, a naked pink, blue, and red neck which ends in a large gular sac, and a huge bill that may be up to 35 cm long. It is widely distributed and common in Africa south of the Sahara, living in various habitats, often near human habitation. Like most storks, it breeds in colonies.

The specific name is derived from the Latin crumena (a purse carried around the neck), naturally referring to the gular sac. The common name is borrowed from French marabout, which was adapted from Portuguese maraboto, which was again derived from Arabic murabit (a soldier stationed in fortified outpost), presumably alluding to the upright stance of this species.

 

 

Marabou stork with inflated gular sack, Virunga National Park, eastern Zaire. (Photo copyright © by Kaj Halberg)

 

 

 

Leptoptilos javanicus Lesser adjutant
This smaller relative of the marabou grows to about 1.2 m tall, with a weight up to 5.5 kg. It has a naked white crown, naked pink and bluish skin around he eyes, and a pink neck with a few ‘punky’ tufts of feathers. It is partial to wetlands, and, unlike the marabou and the greater adjutant (L. dubius), it is not often seen near villages. It is distributed from the Indian Subcontinent eastwards to Vietnam, and thence southwards to Indonesia, with the largest population in Cambodia.

 

 

This damaged lesser adjutant was taken care of at Sepilok Orangutan Rehabilitation Centre in Sabah, Borneo. (Photo copyright © by Kaj Halberg)

 

 

 

Mycteria leucocephala Painted stork
In former days, this pretty bird had a wide distribution in Asia, found in the Indus Valley of Pakistan, and in India, Sri Lanka, Southeast Asia, and southern China. Today, however, it is only common locally in India and Sri Lanka, elsewhere it is either very rare, or has disappeared entirely.

Other pictures, depicting this bird, are found on the pages Silhouettes, and Travel episodes – India 1979: Hunting blackbuck with camera.

 

 

Painted stork, Keoladeo National Park, Rajasthan, India. It is a common breeding bird in this sanctuary. (Photo copyright © by Kaj Halberg)

 

 

 

Columbidae Pigeons and doves
A large family with about 50 genera and c. 345 species. The word pigeon generally denotes larger species, dove smaller species. These birds are found on all continents except Antarctica.

 

Turtur tympanistria Tambourine dove
A small, white and brown dove, which is quite common in woodlands, shrubberies, and plantations, ranging from the Sahel zone and Ethiopia southwards through eastern Africa to south-eastern South Africa. It also occurs on the Comoro Islands.

The generic name is Latin, meaning ‘turtle dove’, whereas the specific name is Ancient Greek, meaning ‘female drummer’, ultimately from tympanon (‘drum’ or ‘tambourine’), alluding to its call, a repeated du-du-du, slightly reminiscent of the sound from a tambourine.

 

 

Tambourine dove, Litipo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Corvidae
Members of this family are found in most parts of the globe. It contains 24 genera with more than 120 species of crows, ravens, rooks, jackdaws, jays, magpies, treepies, choughs, nutcrackers, and many others.

 

Corvus corax Common raven
Divided into at least 8 subspecies, the common raven is the most widespread member of the family, occurring in almost all arctic and temperate areas of the Northern Hemisphere. With a length of up to 63 cm, and weighing up to 1.2 kg, this is the second-largest of passerines, only slightly smaller than the thick-billed raven (C. crassirostris), which lives in Ethiopia. Ravens have been known to live more than 23 years in the wild.

The specific name is a Latinized form of the Greek korax, meaning ‘raven’ or ‘crow’. The English name derives from Old Norse hrafn, ultimately from Proto-Germanic khrabanas.

 

 

Raven chick, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Estrildidae Munias and allies
Munia, mannikin, silverbill, avadavat, waxbill, seedcracker, firefinch, twinspot – members of this family have numerous names. It is large, comprising about 130 species of small, seed-eating birds, distributed across warmer parts of Africa and Asia. Their bill is short and thick, adapted to crushing grass kernels and other seeds.

A large number of these birds are in the pet trade, and many of them have escaped to form feral populations.

 

Hypargos niveoguttatus Red-throated twinspot
A common species, found from Kenya and Zaire southwards to north-eastern South Africa. The male has bright red head and upper breast and black belly with numerous white spots. The female is duller, with grey head and orange or pinkish-red upper breast.

This bird is also known as Peters’s twinspot, described in 1868 by German naturalist and explorer Wilhelm Karl Hartwig Peters (1815-1883) who travelled extensively in southern Africa 1842-1848. He returned to Berlin with an enormous collection of natural history specimens, which he then described in Naturwissenschaftliche Reise nach Mossambique… in den Jahren 1842 bis 1848, ausgeführt (1852-1882), dealing with mammals, birds, reptiles, amphibians, fishes, insects, and plants.

The specific name is derived from the Latin niveus (‘snow-white’) and gutta (‘drop’), of course alluding to the spotted underside.

 

 

Male red-throated twinspot, Pugu Forest, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Falconidae Falcons and caracaras
This family, counting about 60 species, is divided into 3 subfamilies, Herpetotherinae (laughing falcon and forest falcons), Polyborinae (caracaras and Spiziapteryx), and Falconinae (typical falcons and falconets).

 

Falco Typical falcons
The largest genus of the family, comprising about 40 species. It is widely distributed on all continents, except Antarctica.

The generic name is derived from the Latin falx, or falcis (‘sickle’), referring to the claws.

 

Falco rusticolus Gyrfalcon
The largest falcon, females to 65 cm long and weighing up to 2 kg, males smaller, to 60 cm long, weighing up to 1.3 kg. It is a circumpolar breeder, found in Arctic America, Greenland, Iceland, and northern parts of Scandinavia, Finland, Russia, and Siberia, probably southwards to Kamchatka and the Commander Islands. Previously, several races were recognized, but today most authorities regard these as colour morphs.

The specific name is derived from the Latin ruris (‘country’) and colere (‘to dwell’), thus ‘countryside-dweller’. The common name was adapted from French gerfaucon, possibly derived from Old High German gir (‘vulture’), referring to its large size, or from Ancient Greek gyros (‘circle’), alluding to the hunting method of the bird. As opposed to most other falcons, it circles while searching for prey.

 

 

An exhausted gyrfalcon, white morph, resting on board an Icelandic research ship, near Kap Brewster, eastern Greenland. From the cook, I got some bits of meat, tossing them up to the falcon, which ate them greedily. (Photo copyright © by Kaj Halberg)

 

 

 

Falco sparverius American kestrel
This is the smallest falcon in America, growing to about 30 cm long. Divided into about 17 subspecies, it is found in the entire Americas, with the exception of the Arctic, rainforest areas, and the very dry deserts in Chile. It lives in a wide variety of open habitats, including grasslands and farmland, and it has also adapted to urban areas.

 

 

This male American kestrel had been damaged and was taken care of in Tucson Desert Zoo, Arizona. (Photo copyright © by Kaj Halberg)

 

 

 

Gruidae Cranes
Cranes are medium-sized to large birds, comprising 15 species, divided into two subfamilies, Balearicinae with two species of crowned cranes, and Gruinae with 13 species, divided into 4 or 5 genera, depending on authority.

The sandhill crane (Grus canadensis) is described on the page Animals – Birds: Sandhill cranes – a threat to breeding birds.

 

Balearica regulorum Grey crowned crane
Previously, this crane was regarded as being conspecific with the black crowned crane (B. pavonina), but is now regarded as a separate species with two subspecies, regulorum, which is distributed from Zambia and Angola southwards to South Africa, and gibbericeps, which is found from central Kenya and Uganda southwards to Zambia. It is the national bird of Uganda.

The grey crowned crane lives in savannas and river valleys, and also sometimes in agricultural land, provided there are accessible wetlands. Traditionally, some tribes have regarded this bird as sacred, and in many places it has adapted to living near people. It is mainly sedentary, but moves around in search of areas with recent precipitation. It often spends the night in trees.

During the last 50 years or so, this species has declined drastically, and the total population may now be as low as 25-30,000, with two thirds of this number in East Africa. The decline is caused by draining, conversion of savanna to agricultural lands, and lack of flooding due to construction of dams. East African populations are classified as near-threatened, whereas the South African population is stable.

 

 

Feeding grey crowned crane, Ngorongoro Crater, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Grus grus Eurasian or common crane
This is the most widely distributed crane species in the world, found from England eastwards across the entire Siberian taiga to northern Mongolia and China, with small isolated populations in southern Europe, Turkey, the Caucasus, and Tibet. Subspecies archibaldi from Turkey and the Caucasus was described scientifically as late as 2008. Formerly, eastern Asian populations were regarded as a separate subspecies, lilfordi, but today most authorities consider these birds as belonging to the nominate race.

With a total population of nearly 700,000, the Eurasian crane is one of the most numerous crane species, only surpassed by the sandhill crane (G. canadensis). In western Europe, breeding numbers have increased dramatically in several countries, including Sweden with 15,000 to 20,000 pairs, Germany with about 8.000 pairs, and Denmark with about 500 pairs. In 1998, the total number in Europe was estimated at 45,000 pairs, but this number has since increased significantly.

 

 

Eurasian crane, Hornborgasjön, Sweden. This lake is visited by up to 10,000 cranes in March-April, resting here on their way to their breeding grounds. (Photo copyright © by Kaj Halberg)

 

 

 

Laridae Gulls, terns and skimmers
This large cosmopolitan family constitutes 22 genera with about 100 species. A large number of species are described on the page Animals – Birds: Gulls, terns and skimmers.

 

Anous Noddies
A genus of 5 species, distributed in tropical waters.

The generic name is Ancient Greek, meaning ‘stupid’ or ‘foolish’. In the old days, sailors often killed these birds for food, and they found them very stupid, as they were confiding and thus easy to kill.

 

Anous stolidus Brown noddy
This bird is common and widespread, breeding on islets in tropical waters around the world.

The specific name is Latin, meaning ‘stupid’ or ‘foolish’ (see above).

 

 

Brown noddies, nesting on rocks of fossil coral, Latham Island, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Chroicocephalus
This genus with 11 species of small to medium-sized gulls is found almost worldwide. Until recently, they were included in the genus Larus. The generic name is derived from Ancient Greek khroizo (‘to colour’) and kephale (‘head’), alluding to the dark head of many of the species in the breeding season.

 

Chroicocephalus hartlaubii Hartlaub’s gull
This small gull is endemic to the coastline of South Africa and Namibia. Although a coastal species, it rarely ventures far from land. The total population is estimated at about 30,000 birds.

The specific and common names commemorate German physician and ornithologist Karel Johan Gustav Hartlaub (1814-1900), one of the founders of Journal für Ornithologie, in 1852.

 

 

Hartlaub’s gull, Swakopmund, Namibia. (Photo copyright © by Kaj Halberg)

 

 

 

Larus
Formerly, most gulls were placed in this genus, but genetic research has lead to the resurrection of the genera Ichthyaetus, Chroicocephalus, Leucophaeus, and Hydrocoloeus. The systematics of the larger species is very complicated, and a number of former subspecies have recently been elevated to separate species. Today, the genus may contain about 30 species.

 

Larus occidentalis Western gull
This species is common along the Pacific Coast, breeding from Washington southwards to central Baja California, Mexico. Outside the breeding season, it strays to British Columbia and southern Baja California. The largest colony is on the Farallon Islands, west of San Francisco. In the entire San Francisco Bay area, the population is estimated at 30,000 individuals.

It is a large bird, to 68 cm long and weighing up to 1.4 kg. It is divided into two subspecies. The northern nominate race has paler wing colour and darker eye colour than the southern subspecies wymani.

 

 

Incubating western gull, ssp. occidentalis, Yaquina Head, Oregon. (Photo copyright © by Kaj Halberg)

 

 

 

Leiothrichidae Babblers and allies
Most larger babblers are now placed in this family, together with laughing-thrushes of the genera Garrulax, Trochalopteron, Pterorhinus, and Grammatoptila, sibias (Heterophasia), liocichlas (Liocichla), minlas (Minla), barwings (Actinodura), leiothrixes (Leiothrix), and others.

In the past, these genera all belonged to the family Timaliidae, which constituted a true ‘waste-bin of systematics’, in which hundreds of bird species were placed. Today, most of these species have been moved to other families.

 

Argya
A genus of 16 species, distributed across Africa and southern Asia. The majority are medium-sized, long-tailed birds, which forage in noisy groups. They were formerly placed in the genera Turdoides or Garrulax, but following a phylogenetic study in 2018, they were moved to the resurrected genus Argya, which was erected in 1831 by French surgeon and naturalist René Lesson (1794-1849). The generic name is derived from the Latin argutus (‘noisy’).

 

Argya affinis Yellow-billed babbler
This bird, which lives in shrubland, farmland, and larger gardens, is a common resident in South India and Sri Lanka. Nominate subspecies affinis is found in South India, subspecies taprobanus in Sri Lanka.

 

 

Yellow-billed babbler, subspecies affinis, south of Mysore, Karnataka, India. (Photo copyright © by Kaj Halberg)

 

 

 

Monarchidae Monarch-flycatchers
This family contains 16 genera with about 100 species of long-tailed, insectivorous passerines. Most species are sedentary, distributed in sub-Saharan Africa, Southeast Asia, Australasia, and some Pacific islands.

 

Trochocercus African crested flycatchers
Following genetic research, this genus now contains only two members. Three other species, which were previously placed in the genus, have been transferred to the genus Elminia.

The generic name is derived from the Greek trokhos (‘circular’, ’round’) and kerkos (‘tail’).

 

Trochocercus cyanomelas Blue-mantled crested flycatcher
The habitat of this bird is various types of forest. It is found from eastern Zaire, Uganda, and Kenya southwards through eastern Africa to southern South Africa.

The specific name is derived from the Greek kyanos (‘dark-blue’) and melas (‘black’), referring to the bluish sheen in the black plumage of the male.

 

 

Male blue-mantled crested flycatcher, Pugu Forest, near Dar es Salaam, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Muscicapidae Old World flycatchers
A family with about 51 genera and c. 324 species, widely distributed in Africa and Eurasia. Genetic research has had the effect that many smaller birds, which were formerly regarded as belonging to the thrush family (Turdidae), have been moved to this family.

The family name is derived from the Latin musca (‘a fly’) and capere (‘to catch’).

 

Cercotrichas Scrub-robins
Scrub-robins are about 10 species of long-tailed, thrush-like birds, all but one restricted to Africa. The rufous-tailed scrub-robin (C. galactotes) also breeds in southern Europe, the Middle East, and Arabia.

The generic name is derived from Ancient Greek kerkos (‘tail’) and trikhas (‘thrush’), alluding to the long tail.

 

Cercotrichas quadrivirgata Bearded scrub-robin
This species is living in various forest types, scrubland, and gardens, from Kenya southwards to eastern South Africa, westwards to western Zambia and northern Botswana.

The specific name is Latin, meaning ‘four-striped’, referring to the four black stripes on the head, one on the crown, one through the eye, and two moustachial stripes.

 

 

Bearded scrub-robin, Litipo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Cossypha Robin-chats
A genus with about 15 species, living in sub-Saharan Africa.

The generic name is derived from Ancient Greek kossyphos, which was the classical Greek name of the blackbird (Turdus merula), but also meaning ‘thrush’ in general.

 

Cossypha natalensis Red-capped robin-chat
This bright orange-red bird lives in forested areas and dense thickets, from Gabon and Ethiopia southwards to northern Namibia and south-eastern South Africa. It is absent from dry areas and dense rainforest.

The specific name alludes to the KwaZulu-Natal Province in south-eastern South Africa, where the type specimen was presumably collected.

 

 

Red-capped robin-chat, Pugu Forest, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Sheppardia Akalats
A genus of about 10 species of small, robin-like forest birds, found south of the Sahara.

The generic name was given in honour of British farmer Peter A. Sheppard (1875-1958), a settler in Rhodesia and Mozambique. He collected birds for the South African ornithologist Alwin Karl Haagner (1880-1962), who described the east coast akalat (below) in 1909.

The name akalat is of West African origin, used by the Fang and Bulu peoples for members of the genus Illadopsis. However, by mistake, the name was applied to members of Sheppardia in the work The Birds of Kenya Colony and the Uganda Protectorate, by Frederick John Jackson (1860-1929), compiled and edited in 1938 by William Lutley Sclater (1863-1944).

 

Sheppardia gunningi East coast akalat
This skulking bird lives in coastal and montane forests of eastern Africa, from the Sokoke Forest in south-eastern Kenya southwards through eastern Tanzania to Malawi and Mozambique.

The specific name honours Dutch physician Jan Willem Boudewijn Gunning (1860-1913), director of the Staatsmuseum (Museum of Natural History) in Pretoria, and also of the Pretoria Zoological Gardens.

 

 

East coast akalat, Pugu Forest, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Phalacrocoracidae Cormorants
Cormorants and shags are a cosmopolitan family of about 42 species of small to medium-sized, fish-eating birds. Today, most taxonomists divide these birds into three genera: 22 species of mainly larger birds in the genus Phalacrocorax, 15 species of mainly medium-sized birds in the genus Leucocarbo, and 5 species of small birds in the genus Microcarbo. However, the number of genera is disputed, and several new genera have recently been proposed.

Numerous cormorant species are presented on the pages Fishing and Silhouettes.

 

Phalacrocorax
At present, this genus contains 22 species, including the spectacled cormorant (P. perspicillatus), which became extinct in the 19th Century. Several taxonomists propose that many of the species should be transferred to other genera.

The generic name is derived from the Greek phalakros (‘bald’) and korax (‘raven’), where ‘bald’ refers to the white (but feathered) crown of the great cormorant (below) during the breeding season, whereas ‘raven’ presumably alludes to its otherwise black plumage.

 

Phalacrocorax carbo Great cormorant
This bird has an extremely wide, but rather patchy, distribution, found all over Europe and most of Asia, in Australia and New Zealand, and in north-eastern North America and Greenland.

In the 1800s, this species was persecuted all over Europe, partly because it was competing with fishermen, partly because its guano destroyed the trees, in which it was breeding. The complete contrast to this persecution is seen in the Far East, where fishermen, for thousands of years, have been using tamed great cormorants for fishing. Pictures, depicting this practice, are found on the page Fishing.

 

 

Great cormorant, ssp. sinensis, Nature Reserve Vorsø, Horsens Fjord, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Phasianidae Game birds
This family, containing about 190 species, includes pheasants, partridges, junglefowl, Old World quails, peafowl, grouse, and many others. These birds are found worldwide, except in Antarctica.

According to the latest genetic research, guineafowl and New World quails are treated as separate families, Numididae and Odontophoridae, respectively.

 

Gallus gallus Domestic chicken
Recent studies indicate that chickens, or simply fowl, were first domesticated in China about 8000 B.C., descended from the red junglefowl (Gallus gallus), which is still found in the wild in India and Southeast Asia. Later, about 6000 B.C., domestication of the red junglefowl also took place several places in Southeast Asia and in India, where some hybridization with the grey junglefowl (G. sonneratii), living in the southern part of India, may have occurred.

The chicken is described in depth on the page Animals – Animals as servants of Man: Poultry.

 

 

Chick of an old Danish breed, a few days old, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Gallus lafayettii Sri Lankan junglefowl
As its name implies, this bird is restricted to Sri Lanka, where it is widely distributed, living in a wide variety of forested habitats, from sea-level up to an elevation of 2,300 m. It is the national bird of Sri Lanka.

The specific name was given in honour of a French aristocrat, Marie-Joseph Paul Yves Roch Gilbert du Motier, also known as Marquis de Lafayette (1757-1834). He participated in the American Revolutionary War, commanding American troops in several battles. After returning to France, he was a key figure in the French revolutions in 1789 and 1830.

 

 

This gorgeous male Sri Lankan junglefowl was encountered in Sinharaja Forest Reserve. (Photo copyright © by Kaj Halberg)

 

 

 

Picidae Woodpeckers
Members of this family, comprising about 240 species in 35 genera, are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and polar regions. Most species are known for their characteristic way of foraging, pecking on tree trunks and branches. In the breeding season, many species communicate by drumming with their beak on a tree trunk.

 

Dendrocopos
A widespread genus, found in Eurasia and North Africa. Formerly, it contained about 25 species, but many have been moved to other genera, leaving 12 species in Dendrocopos.

The generic name is derived from the Greek dendron (‘tree’) and kopos (‘hard work’), of course alluding to their way of feeding.

 

Dendrocopos major Great spotted woodpecker
Divided into at least 14 subspecies, this bird is very widely distributed, found almost all over Europe, in North Africa and the Middle East, eastwards to the Alborz Mountains of northern Iran, and in a broad belt across the Siberian taiga to Kamchatka, and thence southwards through China, Korea, and Japan to north-eastern India, northern Indochina, and the Island of Hainan. It is mostly resident, although some northern populations spend the winter further south, mingling with the local population.

 

 

This great spotted woodpecker, which was caught in a mistnet on the island of Rømø, Denmark, had been ringed in Norway. (Photo copyright © by Kaj Halberg)

 

 

 

Jynx Wrynecks
A small genus of only two species, the Eurasian wryneck (below) and the red-throated wryneck (J. ruficollis), which lives in sub-Saharan Africa. They got their English name from their ability to turn the head from side to side, and back and forth, writhing in the manner of a snake. This behaviour is thought to imitate a snake, to deter predators from their nest.

The generic name is the Ancient Greek word for wryneck.

 

Jynx torquilla Eurasian wryneck
This small relative of woodpeckers is very widely distributed, found in the major part of Europe, in North Africa, Turkey, and the Caucasus, and in a broad belt across the Siberian taiga to Ussuriland, Sakhalin, northern Japan, and central China. There is also a small isolated population in Kashmir. With the exception of the North African population, it is migratory, spending the winter in the African Sahel zone, in India, Indochina, southern China, and southern Japan. It lives in open areas with scattered trees, including orchards.

 

 

Wryneck, Nature Reserve Vorsø, Horsens Fjord, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Pycnonotidae Bulbuls and allies
These birds are medium-sized passerines, encompassing 27 genera with c. 150 species. They are distributed in much of Africa, and across the Middle East to Tropical Asia and Indonesia, and thence northwards to Japan.

Sub-Saharan members of the family, called greenbuls, brownbuls, leafloves, and bristlebills, are mainly forest birds, whereas the majority of North African and Asian species live in open areas.

 

Bleda Bristlebills
A genus of 5 species, found in rainforests from Senegal eastwards to western Kenya, southwards to northern Angola and southern Zaire. They live in the understorey of forests, where they often follow safari ants on the move, catching insects and other small animals fleeing from the ants.

The generic name alludes to Bleda (died 445), elder brother of Attila the Hun. They ruled their empire together, but Bleda was probably murdered by Attila.

In 1837, when English naturalist and artist William Swainson (1789-1855) described the red-tailed bristlebill (below), he named it Dasycephala syndactyla. However, the name Dasycephala was later applied to a genus in the New World family Tyrannidae. The name of that genus was later changed to Attila, alluding to the aggressive behaviour of members of the genus. Thus we may assume that this is the reason why the red-tailed bristlebill was renamed Bleda. If anyone has a more exact explanation, I would be glad to hear about it.

The common name alludes to the long bristly hairs at the base of the bill.

 

Bleda notatus Yellow-lored bristlebill
This bird, also known as lesser bristlebill, lives in rainforests, from eastern Nigeria and Gabon eastwards through Zaire to Uganda. It was previously regarded as being conspecific with the green-tailed bristlebill (B. eximus), which is today considered to be restricted to West Africa.

Adult birds have a bright yellow iris, younger birds brown.

The specific name is Latin, meaning ‘marked’. It might refer to the eye colour.

 

 

Yellow-lored bristlebill, Bwamba National Park, Uganda. (Photo copyright © by Kaj Halberg)

 

 

 

Chlorocichla
A genus of five forest birds, distributed in Africa south of the Sahel zone, from Sierra Leone eastwards to northern Zaire, southern Kenya, and southern Somalia, southwards to northern Namibia, northern Botswana, and extreme north-eastern South Africa.

Members of this genus are yellowish or pale green. The generic name is derived from Ancient Greek khloros (‘green’) and kikhle (‘thrush’).

 

Chlorocichla flaviventris Yellow-bellied greenbul
This species differs from other members of the genus by living in more open habitats, including dry forests and miombo woodland with scattered trees.

The specific name is derived from the Latin flavus (‘yellow’) and venter (‘belly’), but the entire bird is yellowish, with a conspicuous, narrow, yellow eyebrow.

 

 

Yellow-bellied greenbul, Pugu Forest, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Scolopacidae Sandpipers and allies
A large family with about 15 genera and c. 95 species of waders, found across the globe.

 

Calidris maritima Purple sandpiper
This bird breeds in Arctic mountains in northern Canada, from Banks Island southwards to Ellesmere and Baffin Islands, in Greenland, Iceland, Jan Mayen, Bear Island, Svalbard, along the Scandinavian mountain range, northern Norway, extreme north-western Russia, Franz Josef Land, Novaya and Severnaya Zemlya, and on the Taimyr Peninsula.

It spends the winter months along shores of north-western and western Europe, southwards to Portugal, in extreme southern Greenland, and along the North American Atlantic Coast, southwards to South Carolina.

The specific name is Latin, meaning ‘living at the sea’, derived from mare (‘sea’). In winter, the bird is usually observed along coasts.

 

 

Purple sandpiper in breeding plumage, Rauðanupur, northern Iceland. (Photo copyright © by Kaj Halberg)

 

 

 

Gallinago gallinago Common snipe
The common snipe is the most widespread among 17 members of the genus, found in wetlands in subarctic and temperate regions of Eurasia, eastwards to the Pacific Ocean, southwards to France, Romania, Kashmir, north-eastern China, and Kamchatka. It also breeds on the Azores and the Aleutian Islands. It is found all-year round in western Europe, but the vast majority of the population is migratory, spending the winter in southern Europe, Tropical Africa, the Middle East, the Arabian Peninsula, and Tropical Asia.

There are two subspecies, faeroeensis in Iceland, the Faroes, and the Shetland and Orkney islands, and the nominate gallinago in the rest of the distribution area. Previously, Wilson’s snipe (G. delicata) of North America was also regarded as a subspecies of the common snipe.

The generic and specific names are the Latin word for snipe, derived from gallina (‘hen’) and the suffix –ago (‘resembling’).

 

 

Common snipe, subspecies faeroeensis, Aðaldal, northern Iceland. (Photo copyright © by Kaj Halberg)

 

 

 

Spheniscidae Penguins
These unique flightless seabirds, comprising 6 genera with about 20 species, are breeding along coasts in Antarctica, New Zealand, southern Australia, southern Africa, southern and western South America, and the Galapagos Islands.

 

Spheniscus demersus Jackass penguin
This bird, named for its donkey-like braying, is confined to coasts of South Africa and Namibia. Its numbers have declined drastically since 1800, when the population was estimated at 4 million. Today, there are proably as few as 20,000 – a result of overfishing as well as climate change.

 

 

Jackass penguins in their breeding colony at Boulders, Simonstown, Cape Town, South Africa. The bird in the lower picture is braying. (Photos copyright © by Kaj Halberg)

 

 

 

Strigidae Typical owls
A large cosmopolitan family, comprising 25 genera with about 220 species. These birds are found on all continents, except Antarctica.

 

Bubo Eagle owls
This genus contains about 20 species of large owls, distributed on all continents, except Australia and Antarctica. In America, they are known as horned owls. The generic name is the classical Latin name of the Eurasian eagle-owl (Bubo bubo).

 

Bubo africanus Spotted eagle-owl
One of the smaller members of the genus, widely distributed in the southern half of Africa, from Gabon, Zaire, and Kenya southwards to the Cape Province of South Africa. Previously, birds in the Sahel Zone, and also in the southernmost parts of the Arabian Peninsula, were regarded as subspecies of this bird, but have now been upgraded to separate species.

 

 

Spotted eagle-owl, Cape Town Zoo, South Africa. (Photo copyright © by Kaj Halberg)

 

 

 

Bubo virginianus American great horned owl
This large owl has an enormous distribution, found from Alaska and northern Canada southwards through the entire United States to southern Mexico and Guatemala, and also in two separate areas of South America, from Columbia eastwards to north-eastern Brazil, and south of the Amazon Basin, from eastern Peru eastwards to the Atlantic Ocean, and thence southwards to north-eastern Argentina.

 

 

Captive American great horned owl, Oregon, United States. (Photo copyright © by Kaj Halberg)

 

 

 

Struthionidae Ostrich
This family contains a single genus with one or two species, found in sub-Saharan Africa.

 

Struthio camelus Ostrich
The ostrich is the largest existing bird, some individuals measuring up to 2.7 m in height and weighing up to 150 kg. Today, it is restricted to open areas of sub-Saharan Africa, from the Sahel zone southwards to South Africa. There are 4 subspecies.

The North African ostrich (ssp. camelus) was once widespread in northern Africa, but has disappeared from large parts of its former range. It is today considered critically endangered.

The Somali ostrich (ssp. molybdophanes), which has a bluish neck and thighs, lives in southern Ethiopia, north-eastern Kenya, and Somalia. Some authorities regard it as a separate species, S. molybdophanes. To me, its seem rather odd that a population of a species, in an area with an otherwise continuous distribution, should constitute a separate species.

The Masai ostrich (ssp. massaicus) has pink neck and thighs. It is found from southern Ethiopia and southern Somalia southwards to southern Tanzania.

The South African ostrich (ssp. australis), is widely distributed south of the rivers Zambezi and Cunene. In some places, it is farmed for meat, leather, and feathers.

A fifth subspecies, the Arabian ostrich (ssp. syriacus), which was formerly very common in the Arabian Peninsula, Syria, and Iraq, became extinct around 1966 due to overhunting.

Attempts to reintroduce the species to Israel have failed. In Australia, escaped ostriches have established feral populations.

 

 

South African ostrich, ssp. australis, in captivity, Zimbabwe. (Photos copyright © by Kaj Halberg)

 

 

 

Sturnidae Starlings and mynas
Starlings and mynas constitute a family of about 118 species of medium-sized passerines, divided into c. 30 genera. Their natural area of distribution includes Europe, Africa, Asia, northern Australia, and some Pacific islands.

Many species have been introduced elsewhere, including North America, Hawaii, and New Zealand. They often compete with native birds, and many are regarded as invasive species, in North America especially the common starling (Sturnus vulgaris), which is dealt with on the page Nature: Invasive species.

 

Notopholia corusca Black-bellied starling
This bird, previously known as Lamprotornis corruscus, is now the sole member of a newly erected genus, Notopholia. It mainly lives in coastal forests, from Kenya southwards to southern South Africa.

The scientific name is derived from the Greek notos (‘south’) and pholia (‘living in a cave or hole’), and the Latin coruscus (‘brilliant’, ‘flashing’), thus ‘the brilliant southern one, living in a cave’, i.e. breeding in a hole in a tree.

 

 

The iris of the black-bellied starling is usually yellow, but for a brief period during the breeding season, the iris of the male turns red. – Litipo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Sulidae Gannets, boobies
A family of 10 species of medium-sized to large seabirds, the smaller boobies (Sula and Papasula) living in tropical or subtropical waters, the larger gannets (below) in temperate seas.

 

Morus Gannets
Gannets are large seabirds, comprising 3 species, the northern (M. bassanus), the Cape (below), and the Australian (below). They are all very similar, but are generally regarded as separate species, although some authorities consider the latter two to be subspecies of M. bassanus.

Gannets hunt fish by diving into the sea from heights of up to 30 m, achieving speeds of almost 100 km/h, which enables them to catch fish at a much greater depth than most other airborne birds.

The generic name is derived from the Greek moros (‘foolish’), given due to the lack of fear shown by these birds at the breeding ground, allowing them to be easily killed. The common name comes from Old English ganot, ultimately from Proto-Germanic ganzo (‘gander’), which, in German, means ‘a male goose’, but in English it may also mean a ‘fool’.

 

Morus capensis Cape gannet
This bird nests in large and dense colonies on islands near the coasts of Namibia and South Africa. In 1996, the total population was estimated at about 340,000 birds. Outside the breeding season, they stray far from the breeding area, often observed as far away as the Gulf of Guinea and Mozambique.

 

 

Cape gannets in courtship display, Lambert’s Bay, South Africa. (Photos copyright © by Kaj Halberg)

 

 

 

Morus serrator Australian gannet
This bird was first known to scientists during Captain James Cook’s first great voyage on board the Endeavor (1768-1771), when English naturalist Sir Joseph Banks (1743-1820) shot three specimens on 24th December 1769, off Three Kings Islands, New Zealand.

In his journal, Banks noted: “Calm most of the day: myself in a boat shooting, in which I had good success, killing cheifly several Gannets or Solan Geese, so like European ones they are hardly distinguishable from them. As it was the humour of the ship to keep Christmas in the old fashiond way, it was resolved of them to make a Goose pye for tomorrows dinner.”

On Christmas Day, Banks recorded that “our goose pye was eaten with great approbation.” – Therefore, it is hardly surprising that none of these specimens found their way to England, considering that a goose is best when fried with the skin on!

This species breeds along the coastlines of Australia (Victoria and Tasmania), and New Zealand, mainly on offshore islands, but also in mainland colonies. Outside the breeding season, it may be observed along coasts from western Australia to Queensland, and around New Zealand, Lord Howe Islands, and Norfolk Islands.

 

 

Preening Australian gannet, Muriwai Beach, New Zealand. Another picture of this species is found at the top of this page. (Photo copyright © by Kaj Halberg)

 

 

 

Threskiornithidae Ibises and spoonbills
A worldwide family with 14 genera of long-legged water birds, found on every continent, except Antarctica. Traditionally, this family has been classified into two subfamilies, the ibises and the spoonbills. However, a recent genetic study concludes that spoonbills are closely related to the Old World ibises, whereas the New World ibises are an early offshoot. (Source: J.L. Ramirez, C.Y. Miyaki & S.N. del Lama 2013. Molecular phylogeny of Threskiornithidae (Aves: Pelecaniformes) based on nuclear and mitochondrial DNA. Genetics and Molecular Research 12 (3): 2740-2750)

 

Platalea leucorodia Eurasian spoonbill
This striking bird breeds in a huge area across the central parts of Eurasia, from the Black Sea eastwards to Korea, and also in southern Iraq, Iran, southern Pakistan, India, and Sri Lanka. In Europe, it breeds at scattered locations, northwards to Denmark, and in Africa it is found in Mauritania and along the Red Sea. Northern populations are migratory, spending the winter in southern Europe, northern Africa, and warmer parts of Asia.

The generic name was the Ancient Greek name of the spoonbill, derived from platea (‘broad’), alluding to the spatula-shaped bill. The specific name is derived from the Greek leukos (‘white’) and erodios (‘heron’).

 

 

Eurasian spoonbill with a chick, Keoladeo National Park, Rajasthan, India. (Photo copyright © by Kaj Halberg)

 

 

This picture from Keoladeo National Park clearly shows the spatula-shaped bill of the spoonbill. (Photo copyright © by Kaj Halberg)

 

 

 

Turdidae Thrushes
Many members of this family are described on the page Animals – Birds: Thrushes.

 

Geokichla guttata Spotted ground-thrush

 

 

This spotted ground-thrush has been caught in a mistnet to be ringed, Rondo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Neocossyphus rufus Red-tailed ant-thrush
This species is found in two geographically separate areas, from southern Cameroun and Gabon eastwards across northern Zaire to western Uganda, and in coastal forests of Kenya and Tanzania, including Zanzibar.

The specific name is Latin, meaning ‘rufous’, referring to the predominantly rufous plumage of this bird.

 

 

This red-tailed ant-thrush has been caught in a mistnet to be ringed, Rondo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Reptiles

 

Agamidae Agamas
Agamas are a large group of lizards, comprising 6 subfamilies with about 64 genera and more than 300 species, distributed in Asia, Australia, Africa, and southern Europe.

 

Calotes Forest lizards, garden lizards
This genus of the subfamily Draconinae contains about 29 species, native from the Indian Subcontinent eastwards to southern China, and thence southwards through Indochina and Malaysia to Indonesia. The greateast diversity is found India, Bangladesh, Sri Lanka, and Myanmar.

The generic name is derived from Ancient Greek kalos (‘beautiful’), referring to the beautiful colours of many of the species.

 

Calotes calotes Southern green forest lizard
This large agamid, growing to 65 cm long including the long, slender tail, lives in forests of Sri Lanka and in montane areas of south-western India. It is bright green with white vertical stripes on the body. In the breeding season, or when excited, the male develops a bright red head and throat, and it may also become a much darker green.

 

 

Southern green forest lizard, Sinharaja Forest Reserve, Sri Lanka. (Photo copyright © by Kaj Halberg)

 

 

 

Calotes versicolor Oriental garden lizard
This species is also known as changeable lizard, or bloodsucker, the latter name alluding to the bright red head and throat of the male in the breeding season, or when excited. It is common, living in a wide range of habitats, including urban areas.

It is distributed from eastern Iran and Afghanistan eastwards across the entire Indian Subcontinent to southern China, and thence southwards through Indochina to Sumatra. It has also been introduced to other places, including Oman, Mauritius, the Seychelles, Singapore, and the United States. In Singapore, it is a threat to the native green crested lizard (Bronchocela cristatella).

In China, it is declining due to persecution, presumably collected for food or traditional medicine.

The specific name is derived from the Latin verso (‘to turn’) and color (‘colour’), thus ‘the one that changes its colour’.

 

 

Male oriental garden lizard, displaying a bright red head, Manas National Park, Assam, north-eastern India. (Photo copyright © by Kaj Halberg)

 

 

 

Otocryptis Kangaroo lizards
These small agamids, comprising two species, are endemic to Sri Lanka.

The generic name is derived from Ancient Greek ot-, the stem of ous (‘ear’) and kryptos (‘hidden’), thus ‘with hidden ears’. The common name alludes to the very long legs of these lizards.

 

Otocryptis wiegmanni Brown-patched kangaroo lizard
This species lives in forests of the wet zone and lower mountains of south-western Sri Lanka, up to altitudes around 1,300 m. The male has a large, elongated, brown patch on his dewlap, whereas the very similar O. nigristigma has a black patch. The latter seems to be restricted to the dry zone of the island.

 

 

Brown-patched kangaroo lizard, Sinharaja Forest Reserve, south-western Sri Lanka. (Photo copyright © by Kaj Halberg)

 

 

 

Stellagama stellio Starred agama
Also known by a number of other names, including rough-tailed rock agama, hardun, and painted dragon, the starred agama is found from western and southern Turkey southwards through Syria and north-western Iraq to southern Jordan, north-western Saudi Arabia, and the Sinai Peninsula, on a number of islands along the west coast of Turkey, on Cyprus, in an area of Greek Macedonia, and on the island of Corfu. It has also been introduced to Malta.

This animal, which may attain a length of up to 35 cm, used to be included in the large genus Agama, but is now the single species in the genus Stellagama. However, it may constitute a species complex and may be split into several species in the future. The generic name is from the Latin stella (‘star’), referring to its many spots.

 

 

Male starred agama, Pergamon, Turkey. (Photo copyright © by Kaj Halberg)

 

 

 

Alligatoridae Alligators and caimans
This family contains 8 species of crocodile-like animals, 2 alligators in a single genus and 6 caimans in 3 genera. Alligators live in the United States and China, whereas caimans are restricted to Central America and northern South America.

 

Alligator mississippiensis American alligator
This large animal, males growing to 4.6 m long and weighing up to 450 kg, lives in swamps in south-eastern United States, from Florida, Mississippi, Louisiana, and eastern Texas northwards to Tennessee and North Carolina.

In the 1800s and the first half of the 1900s, hunting and habitat depleted its numbers seriously, and it was feared that it would not survive. In 1967, it was listed as an endangered species and was legally protected. Since then, conservation efforts have been successful, and today it is again common, and has even expanded its range to Tennessee.

 

 

American alligator, Aransas National Wildlife Refuge, Texas. (Photo copyright © by Kaj Halberg)

 

 

 

 

Chamaeleonidae Chameleons
This family of unique reptiles, comprising 12 genera with more than 200 species, is distributed in Africa, Madagascar, southern Spain, Corsica, Sardinia, southern Italy, southern Greece, the Near East, the Arabian Peninsula, southern India, and Sri Lanka. The vast majority of the species are found in Africa south of the Sahara, and in Madagascar.

The eyes of these animals move independently, and they are able to analyze two different images simultaneously when hunting for prey. When a suitable prey has been spotted, the chameleon projects its very long, sticky, folded-up tongue at a tremendous speed at the prey, often from quite a distance. The poor victim sticks to the tongue, which is then folded up and retracted into the mouth, prey and all.

Chameleons are also able to change colour very fast, which mostly happens when they get excited. An example is shown below under Chamaeleo dilepis.

The family name is derived from the Greek khamai (‘on the ground’) and leon (‘lion’), thus ‘earth-lion’, undoubtedly alluding to the hunting method and voracious appetite of these animals.

Several species are popular pets, and some places they have escaped (or have been released) to form feral populations, which are often a threat to the local insect fauna – and thereby indirectly to insectvorous birds, and plants which are dependent on insects for pollunation. This is seen in Hawaii and other places.

 

Chamaeleo
Today, this genus contains about 14 species. Previously, it was much larger, but the major part of the species have been moved to other genera.

 

Chamaeleo africanus African chameleon
Also known as the Sahel chameleon, this animal is found in the Sahel zone, living in savannas from Mauritania eastwards to Sudan, and thence northwards along the Nile to Egypt. It is a large species, growing to 34 cm long, including the tail. It is often green with many black spots and lateral yellow stripes along the side of the body. It has a large bony casque on its head.

At some stage, it was brought from Egypt to Peloponnese, Greece, which today has a small population of about 350 individuals.

 

 

African chameleon, north of Malbaza, Niger. (Photo copyright © by Kaj Halberg)

 

 

 

Chamaeleo anchietae Double-scaled chameleon
This species occurs from Angola and southern Zaire eastwards to western Tanzania.

The specific name honours Portuguese explorer and naturalist José Alberto de Oliveira Anchieta (1832-1897), who travelled extensively in Portuguese Angola between 1866 and 1897, where he collected a large number of animals and plants. He died in Angola, presumably of malaria.

 

 

Double-scaled chameleon, Sao Hill, western Tanzania. (Photos copyright © by Kaj Halberg)

 

 

 

Chamaeleo dilepis Flap-necked chameleon
Divided into 8 subspecies, this chameleon is widespread and common in sub-Saharan Africa, from Cameroun eastwards to Ethiopia and Somalia, and thence southwards to northern South Africa. It lives in a variety of habitats, including savannah, woodland, shrubberies, and bushy grasslands, and it is also often encountered in villages and suburbs.

It is a large species, up to 35 cm long, including the tail. It may be green, yellowish, or brownish, but there is usually pale stripes on the flanks and underside.

The specific name is Ancient Greek, meaning ‘with double scales’, which may refer to the pattern on the gular sac, where rows of green double-scales alternate with rufous bands (see picture below).

The common name alludes to two flaps on the hind neck, similar to those of the African chameleon (above).

 

 

This excited flap-necked chameleon, with expanded gular sack, displays numerous black spots, Masasi, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Kinyongia multituberculata West Usambara two-horned chameleon
As its name implies, this chameleon is endemic to the West Usambara Mountains in northern Tanzania. It lives in forest, scrubland, and along roads, at elevations between 1,200 and 2,500 m.

 

 

West Usambara two-horned chameleon, Usambara Mountains, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Cheloniidae Sea turtles

 

Lepidochelys Ridleys
A small genus of 2 species, the olive ridley (below) and the endangered Kemp’s ridley (L. kempii), which is restricted to the Gulf of Mexico.

The generic name is derived from Ancient Greek lepidos (‘scale’) and chelys (‘turtle’), perhaps alluding to the scaly appearance of the head and flippers of these animals. The common name is of unknown origin.

 

Lepidochelys olivacea Olive ridley
This rather small species, growing to about 60 cm long, is the most abundant of all sea turtles. It occurs in all warmer seas, primarily in the Pacific and Indian Oceans. It is known for its spectacular synchronised mass nesting, called arribada (Spanish for ‘arrival’), where thousands of females gather on the same beach to lay eggs. Two of these places are Playa Ostional, Guanacaste Province, Costa Rica, and Odisha, eastern India.

The specific name alludes to the olive tinge on this otherwise brownish or greyish animal.

My experience with this animal is described on the page Animals – Reptiles and amphibians: Reptiles and amphibians in the Indian Subcontinent.

 

 

This olive ridley has been caught by scientists to be tagged, near the Maipura River delta, Odisha, India. (Photo copyright © by Kaj Halberg)

 

 

 

Crocodylidae Crocodiles
These huge reptiles, altogether 17 species in 3 genera, are widely distributed in tropical areas, with 3 species in Africa and western Madagascar, 9 species in Asia, New Guinea, and northern Australia, and 5 in the Americas.

 

Crocodylus niloticus Nile crocodile
This formidable predator, some specimens growing more than 5 m long and weighing over 400 kg, is able to bring down large animals like zebras and wildebeest. It is widespread throughout sub-Saharan Africa, with the exception of the arid areas of the south-western part of the continent, and is also found along the west coast of Madagascar. In historic times, it was living all along the Nile River (hence its name), northwards to the river delta. In Egypt, it is now restricted to the southernmost parts, around Lake Nasser.

The largest specimens occasionally turn into dangerous man-eaters.

 

 

Nile crocodile in morning light, Lake Baringo, Kenya. (Photo copyright © by Kaj Halberg)

 

 

Nile crocodile in a breeding centre, Zambezi River, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Crocodylus palustris Marsh crocodile, mugger crocodile
This crocodile occurs in the Indian Subcontinent, including Sri Lanka, and also in southern Iran, living in freshwater habitats, preferring slow-moving, shallow waters.

The largest known specimen measured 5.63 m, but on average males reach a length of 3-3.5 m, females 2-2.5 m, males weighing up to 200 kg. Its numbers have decreased alarmingly during the last 50 years due to habitat destruction (dam building and conversion of wetlands to farmland). They occasionally get entangled in fishing nets and drown, and some are killed by fishermen who regard them as competitors. In 2013, it was estimated that less than 8,700 individuals were living in the wild.

The word mugger is in fact pronounced ‘magar’, the Hindi name of this animal, derived from the Sanskrit makara, referring to a mythical crocodile-like creature.

 

 

Marsh crocodile, Kasara Crocodile Breeding Centre, Chitwan National Park, Nepal. (Photo copyright © by Kaj Halberg)

 

 

 

Crocodylus porosus Saltwater crocodile
As its name implies, this species is living in coastal areas, from south-eastern India and Sri Lanka eastwards to Indochina, and thence southwards through Indonesia and the Philippines to New Guinea and surrounding islands, and northern Australia.

It is the largest living reptile, males sometimes growing more than 6 m long and weighing up to 1,300 kg. Females are much smaller, rarely exceeding 3 m in length. The largest specimens occasionally turn into dangerous man-eaters.

 

 

Saltwater crocodiles in captivity, a sub-adult (top), and a juvenile, Port Blair Zoo, Andaman Islands, India. (Photos copyright © by Kaj Halberg)

 

 

 

Crocodylus siamensis Siamese crocodile
This is a fairly small freshwater crocodile, the largest specimens growing to 3.2 m long and weighing 150 kg. It is native to Indochina, Borneo, and possibly Java, but is critically endangered and has already been extirpated from most of its former distribution area. However, it is kept in captivity in several countries for the skin trade, and is not in imminent danger of going extinct.

 

 

Siamese crocodiles in a commercial breeding centre, Singapore. (Photo copyright © by Kaj Halberg)

 

 

 

Elapidae
A huge and diverse family of snakes, comprising 55 genera with about 360 species, including highly venomous groups like mambas, cobras, kraits, and sea snakes. They are characterized by their permanently erect fangs at the front of the mouth. They live in tropical and subtropical regions around the world, with terrestrial forms in Asia, Australia, Africa, and the Americas, and marine forms in the Pacific and Indian Oceans. They vary greatly in size, from the king cobra (Ophiophagus hannah), which may sometimes grow almost 6 m long, to the white-lipped snake (Drysdalia coronoides), which is a mere 18-20 cm long.

 

Dendroaspis angusticeps Eastern green mamba
This species, one out of 4 in the genus, may grow to about 2 m long. It is a highly venomous, arboreal snake, native to south-eastern Africa, mainly coastal areas, found in Kenya and coastal Tanzania, and from Zimbabwe southwards to the east coast of South Africa.

During field work in 1989 in Tanzanian coastal forests, my companions and I encountered green mambas on several occasions, as they lay draped over thin branches in the trees. However, they were not at all aggressive, and we could admire their beautiful colour from a distance.

 

 

Eastern green mamba in captivity, Nairobi Snake Park, Kenya. (Photo copyright © by Kaj Halberg)

 

 

 

Gavialidae
A small family of crocodile-like reptiles, comprising only two members, the gharial (below) and the false gharial (Tomistoma schlegelii), both living in Asia.

 

Gavialis gangeticus Gharial
This animal resembles a crocodile, but its snout is long and slender, with 110 sharp, interlocking teeth – a perfect adaptation for catching fish. Males may grow very large, to 6 m long, whereas females are smaller, to 4.5 m long. It lives in rivers in the northern part of the Indian subcontinent, only leaving the water to bask in the sun, or to dig a nest in a sandy shore.

The generic and common names both refer to a distinct lump that the adult male has at the end of the snout, which resembles a clay pot, known as a ghara. This lump is hollow, amplifying a hissing sound that the male emits, which can then be heard far away. The specific name means ‘of the Ganges (River)’.

The gharial population has declined drastically since the 1930s, and the current number may be less than 1,000 individuals. It is listed as critically endangered on the IUCN Red List.

 

 

Captive gharial, showing its long rows of formidable teeth, Kasara Crocodile Breeding Centre, Chitwan National Park, Nepal. (Photo copyright © by Kaj Halberg)

 

 

Male gharial, basking in the sun, Ramganga River, Corbett National Park, Uttarakhand, India. The lump on the tip of the snout gave rise to its name (see text). (Photo copyright © by Kaj Halberg)

 

 

 

Gerrhosauridae Plated lizards
This family, comprising 7 genera with about 37 species, is native to Africa and Madagascar. They are named for the rectangular scales of many of the species.

 

Matobosaurus validus Giant plated lizard
Previously, this large lizard, growing to 60 cm long, was included in the genus Gerrhosaurus, but was moved to a new genus, Matobosaurus, in 2013. Furthermore, the populations in Namibia and extreme southern Angola were split to form a separate species, M. maltzahni. Today, M. validus is found from extreme southern Zambia, north-eastern Botswana, and Zimbabwe southwards to north-eastern South Africa, living in rocky areas.

The generic name is derived from the Ndebele word matobo, meaning ‘bald head’, referring to the Matobo Hills in southern Zimbabwe, which are characterized by rather smooth granite hills. This area is prime habitat for plated lizards.

 

 

Giant plated lizard, Hwange National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Scincidae Skinks
A huge, almost worldwide family, counting about 1,500 species of small to medium-sized lizards, many with smooth and shiny scales.

 

Trachylepis Striped skinks
A large genus of about 90 species, primarily distributed in Africa. One species, T. atlantica, is found on the Brazilian island of Fernando de Noronha. Its ancestors presumably drifted across the Atlantic on a floating tree, sometime during the last 9 million years. Two other species of the genus may also occur on mainland South America, but they are poorly known.

Members of the genus were formerly included in the genera Mabuya and Euprepis.

The generic name is derived from Ancient Greek trakhys (‘rough’) and lepis (‘scale’), alluding to 3 or more slight longitudinal keels on the dorsal scales.

 

Trachylepis wahlbergii Wahlberg’s striped skink
This species, previously regarded as a subspecies of T. striata, is distributed in Namibia, extreme southern Angola, northern Botswana, north-western Zimbabwe, and extreme southern Zambia.

The specific name was applied in honour of Swedish naturalist and explorer Johan August Wahlberg (1810-1856), who travelled in southern Africa between 1838 and 1856, sending thousands of specimens back to Sweden. He was exploring the Okavango area in Botswana, when he was killed by a wounded elephant.

 

 

Wahlberg’s striped skink, Zambezi River, southern Zambia. (Photo copyright © by Kaj Halberg)

 

 

 

Testudinidae Tortoises
Tortoises, comprising about 18 genera with around 50 living species, are distributed on all continents, except Australia and Antarctica. They vary greatly in size, from the famous Galapagos giant tortoise (Chelonoidis niger), which grows to more than 1.5 m long and weighs up to 400 kilos, to dwarves with a length of less than 10 cm.

 

Aldabrachelys gigantea Aldabra giant tortoise
This gigantic tortoise is the second-largest tortoise species, males growing to about 1.2 m long and weighing up to 250 kg, females generally smaller, about 90 cm long and weighing up to 160 kg. It is the only living member of the genus, distributed on a number of islands in the Seychelles. Two other members, which were found in Madagascar, have gone extinct.

The generic name is composed of Aldabra, and the Ancient Greek word for tortoise, khelys.

In historic times, giant tortoises were found on many of the western Indian Ocean islands, as well as in Madagascar, and fossil records indicate that giant tortoises once occurred on all continents, except Australia and Antarctica. Many of the Indian Ocean species were driven to extinction by European sailors, for whom they were a source of food, and they seemed to be all extinct by 1840, except the Aldabra giant tortoise.

 

 

The Aldabra giant tortoise, ssp. gigantea, is endemic to Aldabra Island. This captive specimen at Lake Baringo, Kenya, is eating grass. (Photo copyright © by Kaj Halberg)

 

 

 

Testudo graeca Greek tortoise
This species, also called spur-thighed tortoise, is widely distributed in southern Europe, northern Africa, and western Asia, found in southern Spain, eastern Romania and Bulgaria, Greece, northern Morocco, Algeria, Tunisia, and Libya, and from Turkey southwards to northern Israel and Jordan, and eastwards to the Caucasus, Iran, and Turkmenistan. No less than c. 20 subspecies have been described.

The Greek tortoise may be distinguished from the quite similar Hermann’s tortoise (T. hermanni) by having large symmetrical markings on the top of the head, large scales on the front legs, and a spur on each thigh.

 

 

Greek tortoise, subspecies ibera, Pergamon, Turkey. (Photo copyright © by Kaj Halberg)

 

 

Greek tortoise, subspecies buxtoni, Mian Kaleh, Caspian Sea, Iran. (Photo copyright © by Kaj Halberg)

 

 

 

Varanidae Monitor lizards
This family is a group of about 80 species of large, carnivorous and frugivorous lizards of the genus Varanus, native to Africa, Asia, and Australia.

The family name is of Semitic origin, meaning ‘dragon’ or ‘lizard beast’. The English name is explained in various ways. Some say it has its origin due to the occasional habit of these animals to stand on their two hind legs, seemingly ‘monitoring’ something. Others say that it arose from an old superstitious belief that they would warn people of the approach of venomous animals.

Previously, the earless monitor lizard (Lanthanotus borneensis) was placed in this family, but has since been moved to a separate family, Lanthanotidae.

 

Varanus albigularis White-throated monitor, rock monitor
This species, divided into 3 subspecies, is widespread in Africa south of the Sahara, found from Zaire, Uganda, southern Ethiopia and southern Somalia southwards to South Africa and Eswatini (Swaziland). It is rather large, weighing up to 17 kg. It lives in open, dry habitats, but avoids harsh desert areas.

The family and common names are adapted from the Ancient Syriac word for these animals, sqinqur.

 

 

This eastern white-throated rock monitor, ssp. microstictus, allows ants to crawl on its face, perhaps to get rid of annoying pests, Tunduru, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

Varanus komodoensis Komodo dragon
The Komodo dragon is the world’s largest lizard, growing to 3.5 m long and weighing up to 135 kilograms. This giant monitor lizard has a very limited distribution, found only on the small Indonesian islands of Komodo, Rintja, and Padar. It is described in depth on the page Travel episodes – Indonesia 1985: Difficult journey to Komodo.

 

 

Komodo dragon, Komodo Island. (Photo copyright © by Kaj Halberg)

 

 

 

Varanus salvator Asian water monitor
As its name implies, this large lizard, growing to about 2 m long, lives close to wet habitats. It is native from Sri Lanka and coastal north-eastern India eastwards to Southeast Asia and thence southwards to the Indonesian archipelago. Usually, people chase these animals away, as they are notorious plunderers of chicken yards, but where left in peace, they often become confiding.

The specific name is Latin, meaning ‘saviour’. Its origin is unexplained, but possibly indicates a religious connection.

 

 

This Asian water monitor is resting at the edge of a pond in a city park, Bangkok, Thailand. (Photo copyright © by Kaj Halberg)

 

 

 

Viperidae Vipers, adders, rattlesnakes
A huge, worldwide family of poisonous snakes, comprising 3 subfamilies, the true or pitless vipers (Viperinae) with 13 genera and about 90 species, the pit vipers (Crotalinae) with 22 genera and about 250 species, and Fea’s vipers (Azemiopinae) with 1 genus and 2 species.

The Latin name of the pit vipers, Crotalinae, is derived from Ancient Greek krotalon (‘castanet’), alluding to the rattle on a rattlesnake’s tail. This subfamily is found in Asia and the Americas. The members, which include rattlesnakes, lanceheads, and Asian pit vipers, are distinguished by the presence of a heat-sensing pit organ, located between the eye and the nostril on both sides of the head.

 

Agkistrodon contortrix Eastern copperhead
This strikingly coloured snake is endemic to south-eastern North America, from Massachusetts southwards to north-eastern Mexico. The generic name is derived from the Greek ankistron (a kind of curved chisel) and odon (‘tooth’), alluding to the curved fangs of this species.

Previously, it was assumed that only one species of copperhead existed in the United States, but it has recently been split into 2 species.

 

 

Eastern copperhead in captivity, Nairobi Snake Park, Kenya. (Photo copyright © by Kaj Halberg)

 

 

 

Tropidolaemus wagleri Wagler’s pit viper
This viper is endemic to Southeast Asia, found from southern Thailand eastwards to southern Vietnam, southwards through the Malaccan Peninsula to Singapore and Sumatra with surrounding islands.

This species is sometimes referred to as the temple viper, because of its abundance around the Temple of the Azure Cloud, a Chinese temple in Bayan Lepas, Penang, Malaysia, in daily speech called the Snake Temple.

The specific name was applied in honour of German herpetologist Johann Georg Wagler (1800-1832). He died young of an accidental self-inflicted gunshot wound while out collecting near Munich.

 

 

Wagler’s pit viper in captivity, Temple of the Azure Cloud, Penang, Malaysia. (Photo copyright © by Kaj Halberg)

 

 

 

Amphibians

 

Brevicipitidae
A small family with 5 genera and about 34 species, distributed in eastern and southern Africa.

 

Breviceps Rain frogs
These frogs, comprising about 20 species, mainly live in southern Africa, with a few species in eastern Africa. Many of the species spend long periods underground, mainly emerging after rain – hence the common name. They are also known as short-headed frogs, reflected in the generic name, from the Latin brevis (‘short’) and ceps (‘headed’), the latter derived from Ancient Greek kephale (‘head’).

Males of these frogs are very much smaller than the females, and their limbs are so short that they are unable to cling with their legs to the female during mating. Instead, they emit a gluey secretion from the skin that allows them to ‘stick’ to the female. (Source: amphibiaweb.org)

 

Breviceps adspersus Common rain frog
The most widespread species of the genus is the common rainfrog, often called bushveld rain frog, found in Angola, northern Namibia, Botswana, south-western Zambia, Zimbabwe, Mozambique, Eswatini (Swaziland), and parts of eastern South Africa.

 

 

Common rain frog, Matobo National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Bufonidae True toads
This large family, comprising about 35 genera, is found worldwide, with the exception of Australia, Madagascar, and the polar regions.

The family name is derived from bufo, the Latin word for toads.

 

Bufo bankorensis Taiwan toad
Also known as Bankor toad, this large toad is endemic to Taiwan, widely distributed from sea level up to an altitude of about 3,000 m. Females, which are larger than males, can reach a length of 20 cm from snout to vent. Genetic research indicates that this species originated from Bufo gargarizans of China. It then dispersed to almost the entire island, resulting in highly diverse western and eastern populations.

 

 

This large Taiwan toad in Malabang National Forest, near Hsinshu, is swallowing a one-foot-long earth worm. Note the poison gland behind the eye. (Photo copyright © by Kaj Halberg)

 

 

 

Bufo bufo Common toad
A common species throughout Europe, except the High Arctic, Iceland, Ireland, and most Mediterranean islands, and thence distributed in a broad belt across temperate Asia, eastwards to Lake Baikal, and also in North Africa, Turkey, and the Caucasus.

 

 

A pair of common toads on their way to a pond to lay eggs, Nature Reserve Vorsø, Horsens Fjord, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Duttaphrynus melanostictus Asian common toad
This large species, growing to about 20 cm long, occurs from the Indian Subcontinent eastwards to southern China and Taiwan, and thence southwards through Indochina and Malaysia to Indonesia. It is found from sea level up to elevations around 1,800 m, mostly encountered in rather open habitats, including urban areas and farmland. They avoid dense forests.

The generic name is composed of the name of Indian herpetologist Sushil Kumar Dutta (born 1952), professor of zoology at North Orissa University, Baripada, India, and Ancient Greek phrynos, the term for a kind of venomous frog.

The specific name is derived from Ancient Greek melanos (‘black’) and stiktos (‘punctured’ or ‘spotted’), referring to the black dots on its face.

 

 

Asian common toad, observed on a road, Wufong, western Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Epidalea calamita Natterjack toad
This toad, which was previously included in the genus Bufo, is restricted to Europe, found from Denmark and southern Sweden eastwards to Estonia and north-western Ukraine, and thence in a broad belt across central and western Europe to Spain and Portugal. It is rare in the United Kingdom and Ireland.

The common name refers to the loud rasping call, made by the males in spring, from natter (‘to chat away’), and jack, a word used to name animals, which are smaller than similar kinds, in this case presumably smaller than the common toad (above).

 

 

Natterjack toad, Nature Reserve Tipperne, Ringkøbing Fjord, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Ranidae True frogs
This huge family, comprising at least 27 genera, is distributed throughout most of the world, occurring on all continents, with the exception of Antarctica.

 

Pelophylax esculentus Edible frog
This widespread and common frog, formerly placed in the genus Rana, is a fertile hybrid between the pool frog (P. lessonae) and the marsh frog (P. ridibundus). It is endemic to Europe, occurring from the northern France eastwards to western Russia, and from Denmark, southern Sweden, and Estonia southwards to northern Italy and Bulgaria. It has also been introduced to various countries, including Norway and the United Kingdom.

The specific name is derived from the Latin esca (‘food’) and ulentus (‘abounding in’), thus ‘most edible’. Like the common name it stems from the usage of this frog as food, particularly in France, where its legs are considered a delicacy.

 

 

Edible frog, Lake Tystrup, Zealand, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Tree frogs
A tree frog is any species of frog that spends most of its life in trees. These animals are highly diverse, divided into at least 5 families. Most species are found in the tropics.

 

 

An unidentified tropical tree frog, sitting on a house beam, Niah National Park, Sarawak, Borneo. (Photo copyright © by Kaj Halberg)

 

 

 

Rhacophoridae

 

Polypedates megacephalus Spot-legged tree frog
This species, also known as brown tree frog, is native from central China southwards to Indochina and north-eastern India, and it has also been introduced to Taiwan. It lives in a variety of habitats, such as forests, grasslands, farmland, streams, and ponds.

The female grows to about 8 cm long, the male to 6 cm. It is quite variable in colour, grey, yellow, or reddish-brown on the back, whitish on the belly. The head is triangular, and the eyes are large. When attacked by a predator , it is able to secrete a fluid, which may be toxic or unpalatable. The eggs are laid in white foam nests in trees or bushes over pools.

The generic name is derived from Ancient Greek polys (‘many’) and pedetes (‘leaper’ or ‘dancer’), from pedan (‘to leap’ or ‘to jump’). The specific name is also Greek, derived from megas (‘large’) and kephale (‘head’).

 

 

A pair of spot-legged tree frogs, observed near a pond at the Daoist temple Tzi Gong Da Foa (‘Drunken Grandfather Big Buddha’), Linnei, western Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Invertebrates

 

Orthoptera Grasshoppers and allies
The order Orthoptera is divided into to suborders, Caelifera (grasshoppers) and Ensifera (crickets, katydids, and others). Both groups are characterized by mouthparts adapted for biting and chewing, hind legs modified for jumping, and organs to produce their characteristic sounds. Ensiferans have much longer, threadlike antennae than the grasshoppers, often longer than their bodies.

These animals are plant-eaters, and some species at times become serious pests, forming huge swarms that consume crops over wide areas. Most species protect themselves from predators by camouflage. If detected, they suddenly jump into the air, many species displaying brilliantly coloured wings to startle the predator. Other species have strong colours as a warning to predators that they are ill-tasting.

 

 

Grasshopper, Matobo National Park, Zimbabwe. (Photo copyright © by Kaj Halberg)

 

 

 

Spiny cricket, Rondo Forest, southern Tanzania. (Photo copyright © by Kaj Halberg)

 

 

 

(Uploaded October 2019)

 

(Latest update November 2023)