Rise and fall of the mighty elephants

 

 

Asian elephant (Elephas maximus), Negombo, Sri Lanka. (Photo copyright © by Kaj Halberg)

 

 

These Asian elephants enjoy being washed by their mahouts (trainers) in the Gandak River, Sonpur, Bihar, northern India. The huge crowd of people in the background are Hindu pilgrims, participating in the annual November pilgrimage to this river. (Photo copyright © by Kaj Halberg)

 

 

This savanna elephant bull (Loxodonta africana) is waving his ears, indicating his annoyance with the presence of our vehicle, Tarangire National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

This young savanna elephant, which was drinking from a stream in Etosha National Park, Namibia, seems to be annoyed by the presence of a springbok (Antidorcas marsupialis). (Photo copyright © by Kaj Halberg)

 

 

 

The first elephant-related animals, Eritherium and Phosphaterium, appeared on Earth during the late Palaeocene, about 60 million years ago. However, they resembled tapirs more than modern elephants. From these ancestors, numerous forms evolved. Altogether c. 161 species are known, belonging to five separate families.

In former times, elephants were distributed throughout Eurasia, Africa, and North America, living in a wide variety of habitats and climates, hot and cold. They also varied enormously in size, from giants more than 4 m tall, to dwarves, which stood only 1-1.5 m tall. The latter evolved on isolated islands, including Malta, Cyprus, and Crete in Europe, and the Channel Islands in America, off the Californian coast.

 

 

Eritherium, the earliest and smallest relative of the elephants, lived about 60 million years ago. (Illustration: Public domain)

 

 

 

Present-day elephants
All of these species, but three, have since become extinct. In North America, the mastodon (Mammut americanum) survived until about 10,000 years ago. Presumably, it was eradicated by hunting tribes, which had migrated here from Siberia during the last Ice Age, when, during several separate periods, a land bridge linked Asia with America, across the Bering Strait. The woolly mammoth (Mammuthus primigenius), which lived in Eurasia, survived even longer in Siberia, the last ones dying out only about 4,000 years ago, on the island of Wrangel.

Two of the three surviving species live in Africa, one in Asia. Formerly, the savanna elephant (Loxodonta africana) was distributed all over the African continent, with the exception of deserts and rainforest areas. Today, it is restricted to savannas and semi-deserts in eastern and southern Africa. The closely related, but smaller forest elephant (Loxodonta cyclotis) lives in rainforests of central and western Africa. Some scientists still consider the savanna elephant and the forest elephant as belonging to the same species.

Formerly, the Asian elephant (Elephas maximus) was distributed across Asia, from Asia Minor in the west to northern China in the east, and southwards into Indonesia. Today, it is restricted to small pockets of forest in India, Nepal, Sri Lanka, the Yunnan Province of southern China, Indochina, and on Sumatra and Borneo.

The origin of the Bornean elephant, subspecies borneensis, which is much smaller than other Asian elephants, remains the subject of debate. Some claim that it is descended from introduced elephants. New research, however, states that it may have been living in Borneo for more than 10,000 years. Some scientists even regard it as a separate species, Elephas borneensis.

 

 

A 60-year old bull savanna elephant may have tusks weighing 60 kilos each. This large bull is feeding in a swamp in the Ngorongoro Crater, Tanzania. The savanna elephant has much larger ears than the forest elephant and the Asian elephant. (Photo copyright © by Kaj Halberg)

 

 

The Asian elephant is somewhat smaller than the savanna elephant, but is still a very large animal. This bull is grazing in Corbett National Park, Uttarakhand, India. Note the secretion on the side of its head (see musth in next section). (Photo copyright © by Kaj Halberg)

 

 

Asian elephant bull, Bardiya National Park, western Nepal. (Photo copyright © by Kaj Halberg)

 

 

 

Social animals
Elephants are huge creatures. The savanna elephant is the largest land animal on Earth, a normal bull weighing 5 to 6 tons, a cow around 3 tons. The heaviest elephant ever recorded was a bull, shot in Angola in 1955, which weighed 10 tons and stood more than 4 m tall across the shoulder. On average, a forest elephant weighs only about half as much as a savanna elephant. The Asian elephant is somewhat smaller than the savanna elephant, measuring to 3 m tall across the shoulder, the bull weighing a maximum of 5 tons.

Elephants are very social, living in herds. Smaller herds are family units, led by a matriarch, which is the oldest cow in the herd, with her daughters and their offspring. Young bulls leave the herd, when they reach puberty, living alone or in loose-knit male groups. Sometimes younger females leave the group to form their own herd. However, they still keep in contact with their maternal herd, and smaller groups often join to form larger herds for shorter or longer periods, occasionally comprising hundreds of animals.

Ovulation of females is normally linked to the season of abundance, i.e. the end of the wet season. In periods of drought and famine they simply do not ovulate. When females are receptive, bulls travel long distances to seek them out.

When bulls are in rut, secretions ooze from a gland on the side of their head, signifying a higher production of male hormones, a condition called musth. During musth, bulls are highly aggressive, and tamed Asian elephants in musth have been known to kill their mahout (trainer), without any apparent reason.

The gestation period is almost two years. A new-born savanna elephant calf weighs about 120 kilos, an Asian about 100 kilos. Six years old, the calf weighs about a ton.

 

 

A herd of savanna elephants, drinking from a stream, Tarangire National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

Even though young bull elephants leave the herd to live a solitary life, they often form small, loose-knit groups, like this one in the Ngorongoro Crater, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

A new-born savanna elephant calf weighs about 120 kilos. These pictures are from Lake Manyara National Park, Tanzania. (Photos copyright © by Kaj Halberg)

 

 

A new-born Asian elephant calf weighs about 100 kilos. This picture was taken in an elephant breeding centre near Chitwan National Park, southern Nepal. (Photo copyright © by Kaj Halberg)

 

 

Elephants love bathing, occasionally diving, using their trunk as a snorkel. These pictures show savanna elephants in the Chobe River, northern Botswana (top), and in Ewaso Nyiro River, Samburu National Park, Kenya. (Photos copyright © by Kaj Halberg)

 

 

 

The trunk
Apart from their great size, elephants have a number of unique characteristics. The trunk is in fact the combined nose and upper lip, which have been elongated. The trunk developed, as the elephants grew larger during the Miocene Period, about 20 million years ago, probably to allow them to survive on vegetation with low nutritional value. Because of the heavy weight of their head, they could not develop a longer neck, so instead the trunk was formed. Among its functions are: up-rooting grass tufts; tearing off bark and twigs of trees; picking fruits; smelling; drinking by sucking up water and sprouting it into the mouth; sucking up mud for a mud-bath or dust for a dust-bath; or even to be used as a snorkel when swimming.

 

 

The trunk has numerous functions, such as up-rooting grass tufts. – Savanna elephant, Tarangire National Park, Tanzania (top), and Asian elephant, Yala National Park, Sri Lanka. (Photos copyright © by Kaj Halberg)

 

 

… smelling. – Savanna elephant, Lake Manyara National Park, Tanzania. (Photo copyright © by Kaj Halberg)

 

 

… drinking by sucking up water and sprouting it into the mouth. – Savanna elephants, Etosha National Park, Namibia (top), and Asian elephant, Bardiya National Park, western Nepal. In the upper picture, springboks (Antidorcas marsupialis) and an Angolan giraffe (Giraffa camelopardalis ssp. angolensis) are also present. (Photos copyright © by Kaj Halberg)

 

 

… sucking up mud for a mud-bath. – Asian elephant, Yala National Park, Sri Lanka. (Photo copyright © by Kaj Halberg)

 

 

… or dust for a dust-bath. – Savanna elephants, Amboseli National Park, Kenya. (Photo copyright © by Kaj Halberg)

 

 

 

The tusks
Another unique characteristic is the tusks. These are two elongated incisors, appearing at an age of about two years and growing throughout the elephant’s life. They consist of a mixture of dentine and calcium salts, called ivory. The bull’s tusks are much larger than the cow’s. A 60-year old bull savanna elephant may have tusks weighing 60 kilos each, whereas the tusks of a cow of similar age only weigh about 9 kilos each. The record is a stunning 130 kilos, and a length of 3.5 m.

In former days, these great tuskers were the prime target of trophy hunters, and today no such animals exist. The Asian bulls have smaller tusks than the savanna elephant, and those of the cows are very small indeed. In the Sri Lankan subspecies, maximus, only 10% of the bulls grow tusks, the cows none at all. This trend has been reinforced by poachers, who select the bulls with the largest tusks.

 

 

The tusks are used for various purposes. Here, young savanna elephants are digging out soil from a river bank to eat minerals. – Ewaso Nyiro River, Samburu National Park, Kenya. (Photo copyright © by Kaj Halberg)

 

 

 

The influence of elephants on their environment
Elephants play a significant role in keeping landscapes open. They often turn over trees to get at the foliage, and they may strip trees of bark, causing them to die, thus creating habitats for various insects, woodpeckers, and other animals.

Elephant dung is often visited by butterflies, which suck moisture and nutrients from it. The dung is also much utilized by dung beetles, or scarabs, which form small balls out of the dung, roll them to some chosen place, lay eggs on them, and bury them. The dung is food for their larvae, which are often eaten by various birds, including hornbills. Dung beetles are described on the page Animals: Animal tracks and traces.

 

 

Elephants, eating bark from a fallen baobab (Adansonia digitata), Tarangire National Park, Tanzania. This magnificent tree is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

Danish biologist Thomas Bregnballe at a hollow baobab, which has been partly destroyed by elephants, Tarangire National Park. (Photo copyright © by Kaj Halberg)

 

 

Not only the bark, but also the wood of this baobab in Hwange National Park, Zimbabwe, has been tremendously frayed by elephants. (Photo copyright © by Kaj Halberg)

 

 

 

Elephant dung often attracts butterflies, which suck nutrients from it. This picture shows butterflies of the genus Dixeia, family Pieridae, Meru National Park, Kenya. (Photo copyright © by Kaj Halberg)

 

 

Dung beetle, making a ball out of elephant dung, after which it will roll it away, bury it and lay eggs in it. – Meru National Park, Kenya. (Photo copyright © by Kaj Halberg)

 

 

Southern yellow-billed hornbills (Tockus leucomelas), feeding on insects in elephant dung, presumably larvae of dung beetles, Krüger National Park, South Africa. Many species of hornbill are described on the page Animals – Birds: Hornbills. (Photo copyright © by Kaj Halberg)

 

 

 

Ahmed of Marsabit
One of the most famous elephants ever was Ahmed – a huge tusker, who lived in Marsabit National Park in northern Kenya. Born around 1919, Ahmed grew to become a giant, with enormous tusks, which earned him the nickname ’The King of Marsabit’.

This elephant became so famous that president Jomo Kenyatta, in 1970, declared him ’a living monument’ – a national treasure protected by presidential decree. To avoid him being shot by poachers, Ahmed was guarded day and night by two appointed rangers, and he lived to die of old age in 1974, about 55 years old.

People always believed that his tusks were among the largest ever recorded, but, as it turned out after his death, they weighed ’only’ c. 67 kilos each – far less than other tusks recorded.

Today, a cast of Ahmed is exhibited in front of the National Museum in Nairobi.

 

 

Ahmed of Marsabit. (Photo: Public domain)

 

 

 

Threatened with extinction
In 1989, the African elephant was listed under Appendix I by the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES), making trade illegal. Appendix II status, which allows restricted trade, exists in Botswana, Namibia, Zimbabwe, and South Africa, and in Botswana, Cameroun, Gabon, Mozambique, Namibia, South Africa, Tanzania, Zambia, and Zimbabwe there are export quotas for elephant trophies.

In 2008, the African elephant was listed as vulnerable by the International Union for Conservation of Nature (IUCN), but, unfortunately, there was no distinction between savanna elephant and forest elephant.

Ivory is much treasured for carving, especially by the Chinese, and this is the main reason for the decline of elephants worldwide. Officially, elephants are fully protected, and trade with ivory is strictly controlled. But in spite of these precautions, all three surviving species are threatened with extinction. In several places, almost all bulls have been shot by poachers. In Kerala in South India, for example, there are now 20 cows for every bull, and there may be too few bulls to mate with all cows.

Another big problem is that these huge animals are great wanderers and need large areas to survive. Everywhere across Asia and Africa, humans have encroached on elephant habitat, causing the elephants to eat crops, hereby becoming great pests. Naturally, people want to protect their crops, and there are many instances of people being killed, when they try to scare away elephants from their fields.

In many cases, it has become necessary to restrict the movements of elephants by erecting electrical fences or digging deep trenches, which the huge animals cannot cross. In some cases, whole herds of elephants have been eradicated, as they had become notorious crop raiders.

In all countries with populations of African elephants they are protected in national parks and game sanctuaries. However, in areas, where this protection is efficient, population densities often become too high, causing the elephants to destroy their own habitat. Previously, large-scale culling took place in these cases, but this practice ceased in 1988, and today numbers are controlled by contraception or translocation. However, most of their range (70%) is outside sanctuaries, and in these areas poaching is prevalent.

During the last 35 years, there has been a steady decline in numbers of African elephants. In 1979, the total population was estimated at 1.3 million, and possibly there were as many as 3 million. By 1989, this number had plummeted to about 609,000, of which c. 214,000 were forest elephants – a number much lower than had previously been thought. From 1977 to 1989, elephant populations declined by 74% in East Africa, and after 1987, the savanna elephant declined by 80% across Africa, from Cameroun to Somalia, and the forest elephant declined by 43%. In later years, this trend has ceased in eastern and southern Africa, and populations have increased by c. 4% per annum. In 2012, IUCN estimated a total of c. 440,000 African elephants (both species).

The current range of the Asian elephant is highly fragmented. In 2008, IUCN listed it as endangered due to a 50% population decline over the past 60-75 years. The total population of wild Asian elephants may now be as low as 40,000-50,000, or maybe even lower, as this number is an estimate. Researchers believe that about half of the population lives in India. In Sri Lanka, their number is about 6,000 – half as many as in the 1800s. However, Sri Lanka still has the highest density of Asian elephants anywhere. Numbers are unknown in Southeast Asia, but only scattered herds exist. Almost everywhere the species is declining, particularly in Southeast Asia. It seems, however, that the population in West Ghats, South India, is increasing.

CITES has listed the Asian elephant under Appendix I, which means that trade in elephants and elephant products is illegal.

 

 

During the annual November market in Sonpur, Bihar, northern India, tamed elephants were often for sale, beautifully decorated with paint. This picture is from 1997. (Photo copyright © by Kaj Halberg)

 

 

 

The lower part of the trunk of this old Taiwan red cypress (Chamaecyparis formosensis) resembles a kneeling elephant. – Alishan, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Elephants and people
The role of elephants in culture and religion is described on the page Animals – Animals in culture and religion – Mammals: Elephantidae.

 

 

 

References
africahunting.com/threads/ahmed-the-elephant-of-marsabit.15201
en.wikipedia.org/wiki/Elephantidae
Blanc, J. 2008. Loxodonta africana. IUCN Red List of Threatened Species, version 2012.2. International Union for Conservation of Nature
Choudhury, A. et al. 2008. Elephas maximus. IUCN Red List of Threatened Species, version 2012.2. International Union for Conservation of Nature
Daniel, J. C. 1998. The Asian Elephant – a natural history. Natraj Publishers

 

 

 

(Uploaded February 2016)

 

(Latest update December 2025)