Plants of Sierra Nevada

 

 

Forest of giant sequoia (Sequoiadendron giganteum), Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

Morning light on a forest of quaking aspen (Populus tremuloides), clad in yellow autumn foliage, Conway Summit. (Photo copyright © by Kaj Halberg)

 

 

Cirrus clouds above the Sierra Nevada, seen from the town of Bishop, Inyo County. (Photo copyright © by Kaj Halberg)

 

 

Coniferous forest around Lake Hume. In the background Mount Goddard (4134 m), named after civil engineer George Henry Goddard (1817-1906), who surveyed the Sierra Nevada during the 1850s. (Photo copyright © by Kaj Halberg)

 

 

 

The Sierra Nevada (‘snowy saw-like mountains’ in Spanish) is a huge mountain range in California and Nevada, varying in width from 75 to 120 km, and stretching over more than 600 km, from the Mojave Desert in the south to the Cascade Range in the north, near the border to Oregon. The vast majority of these mountains is situated in California, although the Carson Range lies primarily in Nevada.

The Sierras harbour three national parks: Yosemite, Sequoia, and Kings Canyon, besides 20 wilderness areas and two national monuments. Prominent features in these mountains include Mount Whitney (4421 m), the highest peak in the lower 48 states, Lake Tahoe, the largest alpine lake in North America, and Yosemite Valley, which is certainly one of the most spectacular valleys on the continent, sculpted by glaciers from ancient granite rocks.

 

 

At 496 km2, Lake Tahoe is the largest alpine lake in North America. This picture shows Fanette Island in Emerald Bay. (Photo copyright © by Kaj Halberg)

 

 

Yellow lichens on a rock face, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Sierra Nevada is criss-crossed by streams, large and small. This picture shows Roaring River, Kings Canyon National Park. (Photo copyright © by Kaj Halberg)

 

 

A number of waterfalls are found in Yosemite National Park, including Vernal Fall (top) and Yosemite Fall, here seen with a rainbow, created by reflections in the vapours from the falling water. (Photo copyright © by Kaj Halberg)

 

 

Barren landscape on a hilltop at the southern outskirts of Sequoia National Forest, near Lake Isabella. (Photo copyright © by Kaj Halberg)

 

 

Kings River, Kings Canyon National Park, with incense cedars (Calocedrus decurrens) growing along the shore. (Photo copyright © by Kaj Halberg)

 

 

Two remarkable peaks in Yosemite National Park, El Capitan (left) and Half Dome. They were sculpted from ancient granite rocks by glaciers. (Photo copyright © by Kaj Halberg)

 

 

 

Half Dome is named after its shape. One side is a sheer rock face, whereas the other three sides are smooth and round, making it appear like a dome, cut in half.

An Ahwahneechee legend relates that, long ago, two travellers, Tissiak and her husband Tokoyee, were quarrelling with each other. He became so enraged that he began beating her, which made her so angry that she hurled her basket of acorns at him. As they stood facing each other, they were turned to stone for their wickedness. The acorn basket (Basket Dome) lies upturned beside Tokoyee (North Dome), and the rock face of Tissiak (Half Dome) is stained with her tears.

 

 

Half Dome in evening light. (Photo copyright © by Kaj Halberg)

 

 

 

Plant communities
Due to the wide variety of ecosystems and habitats in the Sierras, these mountains are home to numerous plant communities. The most important plant zones are mentioned below, with an indication of typical indicator trees, followed by presentation of a number of species within each of these zones, except the alpine zone, which I have not visited.

1) Foothill grasslands, oak woodlands, and chaparral (including Great Basin dry zone on eastern slopes), altitude c. 300-900 m. Indicator species: grey pine (Pinus sabiniana) and blue oak (Quercus douglasii). Chaparral is a community of shrubby plants, adapted to dry summers and moist winters, which is typical of southern California.

2) Lower montane forest (dry south-western part), altitude c. 900-2100 m. Indicator species: ponderosa pine (Pinus ponderosa) and Jeffrey’s pine (P. jeffreyi).

3) Lower montane forest (moist north-eastern part), altitude c. 900-2100 m. Indicator species: ponderosa pine (Pinus ponderosa) and giant sequoia (Sequoiadendron giganteum).

4) Piñon pine – juniper woodland (eastern dry slopes), altitude c. 1500-2100 m. Indicator species: singleleaf piñon (Pinus monophylla) and Utah juniper (Juniperus osteosperma).

5) Upper montane forest, altitude c. 2100-2700 m. Indicator species: red fir (Abies magnifica) and tamarack pine, or Sierra lodgepole pine (Pinus contorta ssp. murrayana).

6) Subalpine zone, altitude c. 2700-2900 m. Indicator species: whitebark pine (Pinus albicaulis).

7) Alpine zone, above c. 2900 m. Indicator species: willow shrubs (Salix spp.).

 

 

Foothill grasslands, oak woodlands, and chaparral, c. 300-900 m

 

Trees

 

Aesculus Horsechestnuts, buckeyes
Horsechestnuts, in America known as buckeyes, are native to temperate areas of the Northern Hemisphere, with one species in Europe, c. 10 in Asia, and 7 in North America.

The generic name is the classical Latin name of the sessile oak (Quercus petraea), possibly derived from aigilops, the classical Greek name of an oak with edible acorns. As horsechestnut fruits are poisonous, it is indeed a bit of a mystery, why Swedish naturalist Carl Linnaeus (1707-1778) named the genus Aesculus. However, it must be pointed out that the meal is edible after boiling the fruits, and it was formerly used as cattle and chicken feed.

The name horsechestnut comes in part from at kestanesi, the Turkish name of the European A. hippocastanum, and the horse-part stems from the usage of the fruits to treat ailments in horses, including excessive wind. The name buckeye stems from an American indigenous tribe, who called the nut hetuck, which means buck-eye, alluding to the markings on the nut, which resembles the eye of a deer.

 

Aesculus californica California buckeye
This tree is widely distributed in California, but is found nowhere else, growing along the coast and at lower elevations of the Sierras and the Cascade Range.

Its large nuts are poisonous. Several indigenous peoples, including Pomo, Yokut, and Luiseño, used to catch fish by strewing pounded nuts into streams, causing the stupefied fish to float on the surface.

Other members of this genus are presented elsewhere, European horsechestnut on the page Plants: Ancient and huge trees, and Indian horsechestnut (A. indica) on the page Plants: Himalayan flora 3.

 

 

Three pictures of California buckeye from Sequoia National Park, showing bare limbs just before foliation (top), bursting leaf buds (centre), and a fallen nut. (Photos copyright © by Kaj Halberg)

 

 

 

Pinus Pines
Pines are a large genus, comprising around 125 species, distributed in arctic, temperate, and subtropical areas of almost the entire Northern Hemisphere. If left in peace, many species are able to grow very old and gnarled.

The generic name is the classical Latin word for pines, maybe originally stemming from Sanskrit pitu (‘sap’ or ‘resin’), alluding to the ample production of resin in this genus.

 

 

Pinus sabiniana Grey pine

 

(…) strange-looking Sabine pine (Pinus Sabiniana), which here forms small groves or is scattered among the blue oaks. The trunk divides at a height at fifteen or twenty feet [4,5-6 m] into two or more stems, outleaning or nearly upright, with many straggling branches and long gray needles, casting but little shade. In general appearance this tree looks more like a palm than a pine. The cones are about six or seven inches [15-17 cm] long, about five [12 cm] in diameter, very heavy, and last long after they fall, so that the ground beneath the trees is covered with them. They make fine resiny, light-giving camp-fires, next to Indian corn the most beautiful fuel I’ve ever seen. The nuts, the Don* tells me, are gathered in large quantities by the Digger Indians for food. They are about as large and hard-shelled as hazelnuts – food and fire fit for the gods from the same fruit.”

 

John Muir (1838-1914), Scottish-American writer and environmentalist, in his book My First Summer in the Sierra, 1911.

 

* Mr. Delaney, a sheep farmer, for whom Muir worked during his stay in the Sierra. He was tall and bony, with a sharply hacked profile like Don Quixote.

 

 

The grey pine, also called California foothill pine or digger pine, is an indicator species of the foothill zone in the Sierras. It is endemic to California, found from sea level up to altitudes around 1,200 m.

The specific name honours English lawyer, naturalist, and horticulturist Joseph Sabine (1770-1837) who had a lifelong interest in natural history, a fellow of the Linnean Society and honorary secretary of the Royal Horticultural Society.

The name grey pine alludes to the grey foliage, whereas digger pine may stem from early settlers, who observed Paiute people digging around these trees for their seeds.

 

 

Grey pine is one of the dominating species in the foothill zone of the Sierras. These pictures, however, were taken in Pinnacles National Park, western California. (Photos copyright © by Kaj Halberg)

 

 

 

Quercus Oak
A huge genus with maybe 500 species, 469 accepted by Kew Gardens. They are native to temperate and subtropical areas of the Northern Hemisphere, southwards to Columbia, northern Africa, Sri Lanka, and Indonesia.

Male flowers are in pendent catkins, females mostly in erect spikes. The fruit is a nut, called an acorn, which is partly enclosed in a cupule (see family above), consisting of overlapping bracts, mostly with free tips.

The generic name is the classical Latin term for oaks. It probably stems from the name of the Lithuanian god of thunder, Perkunas. The word oak is from the Anglo-Saxon ek, in ancient Germanic aik, of uncertain origin and meaning.

 

Quercus douglasii Blue oak
This oak, sometimes called mountain oak or iron oak, is endemic to California. It is an indicator species in the foothill zone of the Sierras, and is also common in the Coast Ranges. It was named for its blue-green foliage.

The specific name was given in honour of Scottish gardener and botanist David Douglas (1799-1834), who explored the North American flora during three expeditions. He introduced Douglas-fir (Pseudotsuga menziesii) and other conifers, especially pine species, as well as a number of bushes and herbs, into British cultivation. In a letter to Sir William Jackson Hooker (1785-1865), director of the Botanical Gardens of Glasgow University, he wrote, “You will begin to think I manufacture pines at my pleasure.”

Douglas died under mysterious circumstances while climbing Mauna Kea in Hawaii in 1834. Apparently, he fell into a pit trap and was possibly crushed by a bull that fell into the same trap. He was last seen at the hut of Englishman Edward ‘Ned’ Gurney, a bullock hunter and escaped convict. Gurney was suspected in Douglas’s death, as Douglas was said to have been carrying more money than Gurney subsequently delivered with the body. (Source: Nisbet, J. 2009. The Collector: David Douglas and the Natural History of the Northwest. Sasquatch Books)

 

 

Grassland with open forest of blue oak, Cache Creek Wilderness Area, central California. (Photos copyright © by Kaj Halberg)

 

 

This dead trunk of a blue oak in Cache Creek Wilderness Area serves as a food cache for an acorn woodpecker (Melanerpes formicivorus), which has drilled innumerable holes in the trunk and stored an acorn in each of them. Other pictures, depicting such caches, are shown on the page Animals: Animal tracks and traces. (Photo copyright © by Kaj Halberg)

 

 

 

Quercus garryana Garry oak
This tree, also called Oregon white oak, has a wide distribution, found from southern British Columbia southwards to southern California, growing from sea level to altitudes around 1,800 m. In the Sierras, a local variety, semota, is a shrub, growing to 5 m tall.

It was named in honour of Nicholas Garry (1782-1856), who was deputy governor of the Hudson Bay Company 1822-35.

 

 

Evening light on garry oak, Sequoia National Forest, near Lake Isabella. Acorns of this species are small and rounded. (Photo copyright © by Kaj Halberg)

 

 

 

Quercus wislizeni Interior live oak
An evergreen oak, widespread in California, and also occurring at scattered locations in northern Baja California. It is very common in the lower Sierras.

The specific name was given in honour of Friedrich Adolph Wislizenus (1810-1889), a German-born physician, explorer, and botanist, who first collected it.

 

 

Acorns of interior live oak are narrow and pointed. – Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Herbs

 

Amsinckia Fiddleneck
This genus, comprising about 13 species of the borage family (Boraginaceae), is named after the inflorescence, which is curled like the head of a fiddle.

The generic name was given as a tribute to German businessman and politician Wilhelm Amsinck (1752-1831), who was a benefactor of the Hamburg Botanical Garden.

 

Amsinckia eastwoodiae Eastwood’s fiddleneck
This species is endemic to California, where it grows in various habitats. In the Sierras, it is restricted to the western lower slopes. It has some of the largest flowers of the genus, to 2 cm long and 1.5 cm wide.

It was named in honour of Canadian botanist Alice Eastwood (1859-1953), who created the botanical collection at the California Academy of Sciences in San Francisco.

 

 

Eastwood’s fiddleneck, Three Rivers, just west of Sequoia National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Hesperoyucca Yucca
A small genus with only 2 species, formerly included in the genus Yucca. They are restricted to California and Arizona, and the Mexican states Baja California Norte, Baja California Sur, Sonora, and Sinaloa.

The generic name is composed of Ancient Greek hesperos (‘western’), and the genus name Yucca.

 

 

Hesperoyucca whipplei Chaparral yucca
This impressive plant is native to southern California and Baja California, mainly growing in dry, rocky soils in sage scrub and chaparral, up to altitudes around 2,500 m.

Its narrow leaves are all basal, to 1 m long, grey-green, stiff, saw-toothed, ending in a sharp point. It only has one inflorescence, a large panicle with hundreds of bell-shaped, white or purplish flowers, situated at the end of a thick stalk, which grows very fast, reaching a height of up to 3 m.

Other common names of this spectacular plant include Our Lord’s candle, in allusion to the huge inflorescence, and Spanish bayonet, which, of course, refers to the needle-sharp tips of the leaves.

The specific name honours Amiel Whipple (1818-1863), head surveyor during construction of the Pacific Railroad.

 

 

Chaparral yucca, Sequoia National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Lupinus Lupines
A large genus in the pea family (Fabaceae), comprising more than 200 species, which are mainly distributed in the Americas, but also some species around the Mediterranean and in North Africa.

According to Collins English Dictionary, the generic name, and with that the common name, stems from the Latin lupinus (‘wolfish’), referring to an old belief that these plants would ravenously exhaust the soil.

 

Lupinus stiversii Harlequin lupine
This pretty plant is endemic to California, where it occurs in several separate mountain ranges.

The specific name was given in honour of army physician Charles Austin Stivers, who first collected it in 1862, near Yosemite. The harlequin part refers to the vivid flower colours of this plant, likened to an Italian Middle Age comic figure, Arlecchino (in English Harlequin), who is characterized by his chequered costume.

 

 

Harlequin lupine, Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Lupinus subvexus Valley lupine
Distributed from Washington southwards to California, this species was first described by American botanist Charles Piper Smith (1877-1955). In the Sierras, it is restricted to low altitudes. Its seeds are reputedly poisonous.

Some authorities consider it synonymous with the widespread chick lupine (L. microcarpus).

 

 

Large growth of valley lupine, near Jackson, west of the Sierras. (Photo copyright © by Kaj Halberg)

 

 

 

Triteleia
This genus, containing about 16 species, is found in western North America, from British Columbia southwards to Baja California and Sinaloa, eastwards to Montana and Colorado.

The generic name is derived from Ancient Greek tri (‘three’) and teleos (‘perfect’), referring to the flower parts, which are in multiples of 3, for instance the 6 petals.

Previously, these plants were included in the family Themidaceae, but were since moved to Brodiaeoideae, a subfamily of the asparagus family (Asparagaceae).

 

Triteleia ixioides Prettyface
Naturally, this plant, which is also known as golden star, got its common names in allusion to its pretty, yellow flowers, which are arranged in a spectacular umbel-like cluster, situated at the end of a stem, which may grow to 80 cm long. It is distributed from south-western Oregon southwards to central California, growing in various habitats.

At least 5 subspecies are known, of which scabra, called foothill prettyface, is mainly restricted to the foothills of the Cascades and the Sierras, growing in open sandy places up to elevations around 2200 m.

The specific name means ‘Ixia-like’, alluding to its (slight) resemblance to Ixia, a genus in the iris family (Iridaceae).

 

 

Foothill prettyface, subspecies scabra, Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Great Basin dry zone on eastern slopes, c. 300-900 m

 

Trees

 

Cercocarpus Mountain mahogany
A genus of bushes and small trees of the rose family (Rosaceae), comprising about 12 species, distributed in the western half of North America, from Washington, Idaho, Montana, and North Dakota southwards to central Mexico. Most species grow in drier habitats.

The flowers are small and white, and the fruit is a one-seeded nut (an achene), with the long, plumelike style still attached. The generic name refers to this plume, derived from Ancient Greek kerkos (‘tail’) and karpos (‘fruit’), thus ‘tailed fruit’. The name mahogany alludes to the hard, dark wood of these trees, but they are entirely unrelated to true mahogany, of the family Meliaceae.

 

Cercocarpus betuloides Birch-leaf mountain mahogany
This small tree, growing to 9 m tall, is native to Oregon, California, Arizona, New Mexico, and Baja California. In the Sierras, it is mainly restricted to chaparral.

The leaves are quite distinctive, entire in the lower half, then toothed to the rounded tip. They somewhat resemble birch leaves (Betula), which gave rise to its specific and popular names.

 

 

Fruiting birch-leaf mountain mahogany, near Junction View, Sequoia National Forest. (Photos copyright © by Kaj Halberg)

 

 

 

Yucca
A genus with about 50 species of shrubs and trees in Agavoideae, a subfamily of Asparagaceae. They are characterized by having a rosette of tough, sword-shaped, evergreen leaves, and large terminal panicles of white flowers. They grow in a wide variety of habitats, found from central Canada southwards to Panama.

The generic name builds on a misunderstanding. It is a variant of yuca, which is supposedly derived from kari’na yuca, a Caribbean name of cassava (Manihot esculenta). The word was applied to these plants in 1753 by Swedish naturalist Carl Linnaeus (1707-1778), who may have confused them with the cassava plant.

 

Yucca brevifolia Joshua tree
This striking yucca is characterized by its many branches. It is a typical plant of the Mojave Desert, in the Sierras growing on the lower, dry, south-eastern slopes.

Its popular name stems from Californian Mormons, who encountered this plant on their way to settle with their fellow soul mates in Utah. To them, the outstretched ‘arms’ of this tree recalled the Biblical Joshua, guiding the Children of Israel to the Land of Canaan.

 

 

Typical landscapes on the dry eastern slopes of Sequoia National Forest, with Joshua trees and yellow rabbitbrush (see below). (Photos copyright © by Kaj Halberg)

 

 

 

Shrubs

 

Chrysothamnus Rabbitbrush
This genus, also known as chamisa, contains 9 shrubs of the composite family (Asteraceae), found in the western parts of Canada and the United States, from British Columbia southwards to California, Arizona, and New Mexico, as far east as South Dakota and Nebraska.

The generic name is derived from Ancient Greek khrysos (‘gold’) and thamnos (‘bush’), referring to the golden-yellow flowers of these plants.

 

Chrysothamnus viscidiflorus Yellow rabbitbrush
Yellow rabbitbrush, also known as sticky rabbitbrush, is the most widespread of the genus, occurring in the same area as given above. It is a typical colonizer of disturbed habitats, such as burned land, landslides, and areas prone to flooding.

Previously, it was utilized medicinally by a variety of indigenous peoples, among the Paiute to treat colds and cough, and among the Hopi for skin problems. Gosiute and Paiute produced chewing gum from latex of the roots, whereas Hopi and Navajo made orange and yellow dyes from the flowers.

The specific name is derived from the Latin viscum (a birdlime, made from the mistletoe), and flos (‘flower’), thus ‘sticky-flowered’.

 

 

Yellow rabbitbrush, observed near Junction View, Sequoia National Forest. (Photo copyright © by Kaj Halberg)

 

 

 

Herbs

 

Datura Thorn-apple
A genus with about 14 species of shrubby, very poisonous herbs of the nightshade family (Solanaceae), native to southern United States, Mexico, and Central America.

The generic name is derived from the Hindi name of these plants, dhatura. The common name alludes to the extremely spiny fruit.

 

Datura innoxia Downy thorn-apple
This shrubby herb, also called recurved thorn-apple, prickly-burr, Indian-apple, or moonflower, grows to 1.5 m tall. The most common name stems from the short, greyish hairs, which cover stem and leaves. When bruised, the entire plant has an unpleasant smell of rancid peanut butter. The spectacular white flowers are very large, trumpet-shaped, to almost 20 cm long, in fruit turning into an egg-shaped, spiny capsule, about 5 cm long. The spines often get caught in the fur of animals, which then spread the seeds.

It is native from the south-western United States southwards to Guatemala, and has also been widely introduced as an ornamental, or inadvertently, to all continents except Antarctica.

It contains the highly toxic alkaloids atropine, scopolamine, and hyoscyamine. The Aztecs used this plant long before the Spanish conquest for many therapeutic purposes, including as a poultice for wounds. The Aztecs warned against ingestion of it, as it might cause madness. Nevertheless, various indigenous tribes used it as a hallucinogenic.

The effects of scopolamine and hyoscyamine are described on the page Plants: Plants in folklore and poetry, under Hyoscyamus niger. On the same page, you may read an amusing anecdote from 1676, describing the altered mental state of British soldiers, who consumed common thorn-apple (D. stramonium).

When Scottish botanist Philip Miller (1691-1771) first described this plant in 1768, he misspelled the Latin word innoxia (‘inoffensive’), when naming it Datura inoxia. Another name, D. meteloides, has also erroneously been applied to it.

 

 

Downy thorn-apple, Kern Valley, eastern California. The yellow shrub in the background is yellow rabbitbrush (Chrysothamnus viscidiflorus) (see above). (Photo copyright © by Kaj Halberg)

 

 

Spiral-shaped flower-bud of downy thorn-apple, Sequoia National Forest. (Photo copyright © by Kaj Halberg)

 

 

 

Diplacus Monkey-flower
This genus contains about 50 species, native to western North America, from Washington, Idaho, and Montana southwards to Baja California and Sinaloa.

The generic name is derived from Ancient Greek di (‘double’) and plakos (‘a flat, round plate’). In his publication from 1838, On Two New Genera of California Plants, English naturalist Thomas Nuttall (1786-1859) says: “The generic name alludes to the splitting of the capsule, attached to each valve of which is seen a large placenta, and under its edges are found the slender, subulate seeds.”

These plants were previously included in the genus Mimulus.

 

Diplacus bigelovii Bigelow’s monkey-flower
Formerly known as Mimulus bigelovii, this plant grows in deserts and other dry areas in the south-western United States. It is very variable in size and shape, growing from 2 to 25 cm tall. The flower colour is generally magenta, with a yellow throat.

It is one among many species, which were named in honour of surgeon and botanist John Milton Bigelow (1804-1878), who collected many new plant species on expeditions to south-western U.S. and northern Mexico.

 

 

Bigelow’s monkey-flower, photographed in Joshua Tree National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Eriogonum American buckwheat
A huge genus, comprising maybe 250 species, all occurring in North America and Mexico.

The generic name is derived from Ancient Greek erion (‘wool’) and gony (‘knee’), alluding to the hairy nodes of the first species to be described, E. tomentosum.

 

Eriogonum fasciculatum California buckwheat
This shrubby herb, also known as flat-topped buckwheat, is distributed from central California, Nevada, and Utah southwards to southern Baja California and Sinaloa in Mexico, growing in a variety of habitats, including chaparral, grasslands, sagebrush scrub, and pinyon-juniper woodland.

The stems, to about 1.5 m tall, are spreading or erect, stem leaves singly or in bundles at each node, blade linear or oblanceolate, to 1.5 cm long and 5 mm wide, white-woolly or sometimes almost smooth above, greyish or green below, margin often inrolled. The inflorescence is widely branched, rounded or occasionally flat-topped, with numerous compact heads, to 2 cm across, flowers white or pinkish, to 3 mm across.

In former days, this species was widely used medicinally by various tribes for numerous ailments, including colds, headache, diarrhoea, and wounds. It was also eaten raw or used in porridges and baked items, and tea was made from leaves, stems, and root. Today, it is a very important source of honey.

The specific name is a diminutive of the Latin fascis (‘bundle’), referring to the leaf clusters.

 

 

Fruiting California buckwheat in evening light, Lake Isabella, Sequoia National Forest, California. (Photos copyright © by Kaj Halberg)

 

 

 

Lower montane forest (dry south-western part), c. 900-2100 m

 

Trees

 

Calocedrus Incense-cedars
These conifers of the cypress family (Cupressaceae) are found in two separate areas, with 3 species in the Far East, and one in western North America.

The generic name is composed of Ancient Greek kalos (‘beautiful’), and the genus name Cedrus (‘cedar’). However, these trees are not closely related to true cedars, which belong to the pine family (Pinaceae).

The common name alludes to the leaves, which emit a fragrance when crushed.

 

Calocedrus decurrens California incense-cedar
Formerly called Libocedrus decurrens, this large conifer, growing to about 57 m tall, may have a trunk diameter up to 3 m. Before settlers started logging at a large scale, trees reportedly reached 69 m, with a diameter of 4 m. It grows very slowly, but can reach an age of more than 500 years.

Indigenous peoples utilized it for numerous purposes, the bark to construct temporary conical-shaped huts and also more permanent houses, and to produce fire by friction. Hunting bows and baskets were made from thin branches. It was also used medicinally for stomach problems.

The specific name is Latin, meaning ‘extending down’, referring to the leaves extending down the twig.

 

 

Bark of incense-cedar is thick, on older trees up to 15 cm, with deep furrows, exfoliating in long strips. Yellow lichens are growing on the trunk of this old tree in Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Fruits of incense-cedar, with Liberty Cap in the background, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Dried out trunk of incense-cedar with holes from insect larvae, and fallen needles of ponderosa pine (see below), Kings Canyon National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Pinus lambertiana Sugar pine
A typical tree of this zone, the tallest and most massive of all pines, often growing to 60 m tall, the record being 83 m, and with a trunk diameter to 2.5 m, in some specimens even 3.5 m. It also has the longest cones of any conifer, often growing to 50 cm long, rarely to 65 cm.

It is native from Oregon south through California to Baja California.

Scottish botanist David Douglas (1799-1834) named it lambertiana in honour of English botanist Aylmer Bourke Lambert (1761-1842). The common name stems from its sweet resin, which indigenous tribes used as a sweetener. The famous Scottish-American writer and environmentalist John Muir (1838-1914) preferred it to maple sugar.

The sad fate of David Douglas is related above (see Quercus douglasii).

 

 

Sugar pine is the tallest of all pines, often growing to 60 m high. This towering specimen was photographed in Mariposa Grove, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Branches of a sugar pine, outlined against pale foliage, Mariposa Grove. (Photo copyright © by Kaj Halberg)

 

 

This Douglas’ squirrel (Tamiasciurus douglasii) in Yosemite National Park is feeding on a sugar pine cone larger than itself. This species, and many other squirrels, are described on the page Animals – Mammals: Squirrels. (Photo copyright © by Kaj Halberg)

 

 

 

Pinus ponderosa Ponderosa pine
Also known as bull pine or western yellow-pine, this is an indicator tree of this zone. It grows to a huge size, the record being 81 m. The bark of younger trees up to about 150 years old is very dark, almost black, which later changes into reddish-brown or golden-brown scaly plates, separated by dark furrows.

Ponderosa pine is native to a huge area in western United States and Canada, found from British Columbia southwards to southern California, eastwards to South Dakota and New Mexico.

It was first collected in 1826 by David Douglas (see Quercus douglasii above), who labelled it Pinus ponderosa, from the Latin ponderosus (‘heavy’), referring to its heavy wood.

 

 

Grassland with ponderosa pines, Kings Canyon National Park. (Photo copyright © by Kaj Halberg)

 

 

Ponderosa pine grows very tall, the record being 81 m. These were observed in Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

Ponderosa pine in morning light, near Junction View, Sequoia National Forest. The plants in front of the withered grass are prickly lettuce (Lactuca serriola), an introduced plant from Europe. (Photo copyright © by Kaj Halberg)

 

 

The characteristic bark of ponderosa pine, Kings Canyon National Park (top), and Mariposa Grove, Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

Sprouted ponderosa pine in front of a burned log, Mariposa Grove. (Photo copyright © by Kaj Halberg)

 

 

 

Quercus kelloggii California black oak
This is mainly a tree of northern California and western Oregon, where it grows in foothills and lower mountains. Its occurrence in southern California and Baja California is patchy, but it is common in the Sierras. Most trees live between 100 and 200 years, but some specimens are known to be almost 500 years old.

This species is adapted to fire, protected from smaller fires by its thick bark. It is killed by larger fires, but easily sprouts again from the roots. Acorns mainly sprout, when a fire has cleared an area of leaf litter. This was known by several indigenous peoples, who purposely lit fires to renew growths of this tree, whose acorns was a staple food source to them.

The specific name honours American physician and botanist Albert Kellogg (1813-1887), one of the founding members of the California Academy of Sciences.

 

 

Bark of an old California black oak, Kings Canyon National Park. (Photo copyright © by Kaj Halberg)

 

 

Autumn foliage of California black oak, Kings Canyon National Park. Leaves of this species are deeply cleft, 10-20 cm long. (Photo copyright © by Kaj Halberg)

 

 

Feeding tunnels of a species of leaf miner moth create patterns in a leaf of California black oak, Junction View, Sequoia National Forest. (Photo copyright © by Kaj Halberg)

 

 

Acorns of California black oak are relatively large, about 3 cm long and 1.5 cm wide. – Junction View. (Photo copyright © by Kaj Halberg)

 

 

 

Herbs

 

Anaphalis Pearly everlasting
The majority of these herbs of the daisy family (Asteraceae), comprising about 110 species, are distributed in tropical and subtropical Asia, with only relatively few in temperate areas of Asia, Europe, and North America.

Members of this genus have no ray florets, but very conspicuous, floret-like involucral bracts.

The generic name is derived from Ancient Greek ana, which has several meanings, in this connection probably ‘exceedingly’, and phalos (‘white’), referring to the white, pearl-like flowerheads.

 

Anaphalis margaritacea Common pearly everlasting
Also known as western pearly everlasting, this erect herb grows to 1.2 m tall. It is widespread in North America, from north-western Mexico to Alaska, and in northern Asia, southwards to China and the Himalaya, and thence westwards to eastern Europe.

The specific name is derived from Ancient Greek margarites (‘pearl’), likewise alluding to the white, pearl-like flowerheads.

 

 

In this picture from Kings Canyon National Park, common pearly everlasting has invaded a burned forest of oak and pines. (Photo copyright © by Kaj Halberg)

 

 

Common pearly everlasting, near Junction View, Sequoia National Forest. (Photo copyright © by Kaj Halberg)

 

 

 

Claytonia Spring beauty
A genus of 27 species, which were formerly included in the purslane family (Portulacaceae), but have since been transferred to a separate family, Montiaceae. These plants are distributed mainly in montane areas of northern and central Asia, and in North and Central America.

The genus was named to commemorate botanist John Clayton (c. 1694-1773), who served as an Anglican minister in the Colony of Virginia.

 

Claytonia perfoliata Miner’s lettuce
Formerly known under the names winter purslane and Montia perfoliata, this plant is usually prostrate, but may sometimes grow to 40 cm tall. One of its main characters is the rounded, fleshy leaves, which often encircle the stem.

It occurs from southern Alaska southwards through western United States and Mexico to Central America. It is common in the Sierras.

In Europe, where it was cultivated as a vegetable as far back as in the 1500s, it is widely naturalized today.

The specific name refers to plants, in which the leaves are clasping the stem. The name miner’s lettuce stems from the 1849 Gold Rush in California, when miners ate this plant as a fresh salad, presumably to avoid getting scurvy.

 

 

Miner’s lettuce, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Close-up of miner’s lettuce, Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Epilobium Willow-herb
About 200-220 species of herbs of the evening-primrose family (Onagraceae), distributed almost globally. Many species are glandular-hairy. The fruit is a long capsule, splitting longitudinally, and the seeds have long hairs, an adaptation for wind-spreading.

The generic name was used as early as 1561 for E. angustifolium (today called Chamaenerion angustifolium) by Swiss physician and naturalist Conrad Gessner (1516-1565) in his unfinished work Historia plantarum. He composed the name from Ancient Greek ion epi lobion (‘violet on a pod’), lobion literally meaning ‘fruit of the cowpea (Vigna unguiculata), alluding to the similarity of the flower colour to that of certain violets (Viola), and to its pod-like fruit.

The common name refers to the similarity of the leaves of some species to those of certain species of willow (Salix).

 

Epilobium canum California fuchsia
This shrubby herb, which grows to 60 cm tall, is native from Oregon, Idaho, and Wyoming southwards to northern Mexico, growing on dry slopes and in chaparral.

The specific name is Latin, meaning ‘grey’ or ‘ash-coloured’, presumably referring to the leaves, which may sometimes be greyish, but are usually green.

Previously, it was called Zauschneria californica, named in honour of physician and botanist Johann Zauschner (1737-1799) of Prague. However, genetic research has revealed that it is in fact a species of willow-herb.

The popular name refers to the scarlet flowers, which resemble those of fuchsias. Hummingbirds are much attracted to these flowers, giving rise to two other common names, hummingbird flower and hummingbird trumpet.

 

 

California fuchsia, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Erysimum Wallflower
A genus of about 180 species, belonging to the mustard family (Brassicaceae). These plants are native to Europe, the major part of Asia, and North America, southwards to Costa Rica.

The generic name is a Latinized version of erysimon, the classical Greek name of a species of hedge-mustard, Sisymbrium polyceratium. Some authorities suggest that it stems from erymai (‘to protect’ or ‘to save’), presumably alluding to the usage of Sisymbrium species to cure inflammation of the throat. Why the name was applied to these plants is not clear.

 

Erysimum capitatum Western wallflower
This plant, also called sanddune wallflower or Douglas’s wallflower, has attractive flowers, whose colour varies from bright golden to yellow, red, white, or purple. It is found in western North America, from Alaska southwards to north-western Mexico.

Formerly, it was used medicinally by indigenous peoples for various ailments, including stomach ache and muscle pain.

The specific name is Latin, meaning ‘forming a head’, referring to the dense inflorescence.

 

 

Western wallflower, Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Lower montane forest (moist north-eastern part), c. 900-2100 m

 

Trees

 

Abies Fir
This genus, comprising about 50 species, is mainly distributed in montane areas in central and southern Europe, North Africa, temperate areas of Asia, and much of North and Central America.

The generic name is the classical Latin name of the silver fir (A. alba), probably derived from the Ancient Greek name of the plant, abin. The English name is probably derived from Old Norse fyri, which, however, was the word for pine trees (Pinus).

 

Abies concolor White fir
This impressive tree is native to montane areas of western North America, found at elevations between 900 and 3,400 m. Two subspecies have been described, of which the nominate concolor is found from Wyoming westwards to eastern Nevada, southwards to Arizona and New Mexico, just extending into Mexico, whereas subspecies lowiana occurs from southern Oregon through California to northern Baja California.

White fir may live up to 300 years. The largest specimens have been found in the central Sierras, of which the largest was 75 m tall, with a trunk diameter of 4.6 m at breast height.

The specific name is Latin, meaning ‘of the same colour’, alluding to the needles, which have the same colour above and below.

 

 

Mixed forest, partly burned, of white fir and ponderosa pine (see above), Mariposa Grove, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

White firs, Sequoia National Park. A species of wolf lichen (see below) is growing on the trunks. (Photo copyright © by Kaj Halberg)

 

 

 

Acer Maple
In former days, maples constituted a separate family, Aceraceae, which is now regarded as belonging to Hippocastanoideae, a subfamily of the soapberry family (Sapindaceae).

These plants, comprising about 130 species of deciduous trees or large shrubs, are widespread in the Northern Hemisphere, with most species in Asia, others in Europe, northern Africa, and North and Central America. Only a single species occurs in the Southern Hemisphere.

The leaves are palmate in most species, and the foliage often turns a brilliant red or yellow in autumn. A number of pictures, depicting this autumn foliage, are found on the page Autumn.

The fruit consists of 2 connected single-seeded units, each with a long wing. When ripe, the fruits are often propelled a considerable distance by the wind. An account of the effectiveness of this spreading is described on the page Nature reserve Vorsø: Expanding wilderness.

The generic name is the classical Latin word for maples.

 

Acer macrophyllum Bigleaf maple
As its name implies, this tree, also called Oregon maple, has large leaves – in fact the largest leaves of any maple, often to 30 cm across, with five deeply indented lobes. Usually, it grows 15-20 m tall, but specimens up to 48 m have been encountered. It is native to the Pacific Coast, from southernmost Alaska to southern California. Inland, it occurs in the Sierras, and in central Idaho.

Pictures, depicting the yellow autumn foliage of this species, may be seen on the page Autumn.

 

 

Leaves and fruits of bigleaf maple, Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Alnus Alder
This genus, comprising about 40 species of shrubs or trees, is distributed throughout the northern temperate zone, with a few species in Central America and the Andes. The fruits are very distinctive, woody, resembling diminutive cones.

The generic name is the classical Latin name of alders. Some authorities connect the name with High German elo (’greyish-yellow’), and with Sanskrit aruna (’reddish-yellow’), referring to the fact that the wood of common alder (A. glutinosa), when cut, assumes a bright reddish-brown colour. The common name evolved from the Old English word for these trees, alor, which in turn derived from Proto-Germanic aliso.

 

Alnus rhombifolia White alder
A deciduous tree, to 25 m tall, which grows in moist habitats at altitudes up to around 2,400 m. It is distributed from southern Washington and Idaho through Oregon to southern California.

The specific name refers to the leaves, whose shape is sometimes rhombic.

 

 

Trunk of white alder, covered in grey and orange lichens, Kings Canyon National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Cornus Dogwood
This genus contains between 30 and 60 species, depending on authority. Most are deciduous trees, shrubs, or dwarf shrubs. A few are evergreen. They are native to temperate areas of Eurasia and North America, with China, Japan, and the south-eastern United States as core areas. The fruit is a drupe, which is edible in some species. The taste, however, is quite insipid.

The generic name is Latin, meaning ‘horn’. It was the classical name of Cornelian dogwood (C. mas), whose wood is so dense that it will sink in water. The prefix dog may imply that the fruits of common dogwood (C. sanguinea) are of little value, due to their bitter taste. Alternatively, it may refer to the former usage of dogwood shoots, which were sharpened and used by farmers as cattle prods, called dags. Skewers were also made from the tough and durable wood.

 

Cornus nuttallii Pacific dogwood
This tree, growing to about 25 m tall, has very characteristic inflorescences, with tiny greenish-white flowers clustered in a dense head, which is surrounded by large white bracts. The fruit is a large pink ‘berry’, to 3 cm across, containing 50-100 seeds. It is edible, though not very tasty.

It is native along the Pacific Ocean, from southern British Columbia to southern California. Inland populations are found in the Sierras, and also in central Idaho. In 1956, this tree was appointed as the provincial flower of British Columbia.

In his book My First Summer in the Sierra, from 1911, Scottish-American writer and environmentalist John Muir (1838-1914) writes about this tree: “Nuttall’s flowering dogwood makes a fine show when in bloom. The whole tree is then snowy white. The involucres are six to eight inches [15-20 cm] wide. Along the streams it is a good-sized tree thirty feet [9 m] high, with abroad head when not crowded by companions. Its showy involucres attract a crowd of moths, butterflies, and other winged people about it for their own, and, I suppose, the tree’s advantage.”

The specific name was given in honour of Thomas Nuttall (1786-1859), a British printer, who came to the United States in 1808. Shortly after his arrival, he met botanist Benjamin Barton (1766-1815), who induced a strong interest in natural history in him. During the following years, until 1841, Nuttall undertook several expeditions in America, and numerous plants and animals are named after him.

 

 

Pacific dogwood is easily identified by its large white bracts. – Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Quercus chrysolepis Canyon live oak
This evergreen oak is common from southwestern Oregon through California to northern Baja California, with scattered populations in Nevada, Arizona, New Mexico, and Chihuahua, Mexico. It is the most widely distributed oak in California.

The leathery leaves are entire or toothed, glossy green above. Young leaves are covered in yellowish down beneath, often turning grey and almost hairless the second year. The acorns vary quite a lot, but are mostly ovoid, with a shallow, turban-like, scaly cup, densely covered in yellowish hairs, which gave rise to the specific name, derived from Ancient Greek khrysos (‘gold’) and lepis (‘scale’).

After leaching of the tannins, the acorns were a staple food of many indigenous tribes. They have also been used as a coffee substitute.

 

 

Acorns of canyon live oak, Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Sequoiadendron giganteum Giant sequoia

 

A grove of giant redwood or sequoias should be kept, just as we keep a great and beautiful cathedral.”

 

Theodore Roosevelt (1858-1919), American president 1901-1909.

 

In the wild, this magnificent tree, the only member of the genus, is found only in the Sierras. Some specimens are the heaviest living beings on the planet, the largest ones having an estimated weight of about 2,100 tons.

The following quote of Scottish-American writer and environmentalist John Muir (1838-1914), from 1870, is cited in the book John Muir: His Life and Letters and Other Writings, edited by T. Gifford, Mountaineers Books, 1996: “Do behold the King in his glory, King Sequoia! Behold! Behold! seems all I can say. Some time ago I left all for Sequoia and have been and am at his feet, fasting and praying for light, for is he not the greatest light in the woods, in the world? Where are such columns of sunshine, tangible, accessible, terrestrialized?”

Most people of that period did not share Muir’s enthusiasm. Despite the fact that their wood is fibrous and brittle, and of little use for construction, thousands of these magnificent trees were ruthlessly cut down, between the 1880s and 1924, even though their commercial value was marginal. The heavy trees would often shatter when they hit the ground, and it has been estimated that as little as 50% of the timber came to use. The wood was utilized mainly for shingles and fence posts, or even for matchsticks. – Imagine! From grand tree to matchstick!

Today, fortunately, felling of this species is strictly forbidden, and a few magnificent groves have been saved.

The generic name is composed of the genus name Sequoia (Californian redwood), and Ancient Greek dendron (‘tree’).

Other pictures, depicting these impressive trees, may be seen on the page Plants: Ancient and huge trees.

 

 

The largest known giant sequoia, called General Sherman, grows in Sequoia National Park. It is about 2,300 years old, 84 m tall, with a diameter of 11 m at ground level, a mass of c. 1,500 m3, and an estimated weight of 2,100 tonnes. Note the burned parts on the lower trunk. Due to their thick, spongy bark, these trees are able to withstand forest fires. (Photo copyright © by Kaj Halberg)

 

 

A majestic growth of giant sequoias, Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

People appear like midgets, when they stand next to a giant sequoia. – Mariposa Grove, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Turned-up roots of a giant sequoia, covered in yellow lichens and green mosses, Mariposa Grove, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Umbellularia californica California bay laurel
Mainly coastal, this evergreen tree occurs from south-western Oregon southwards through California to the Mexican border. It is quite common in the western foothills of the Sierras. It is the only member of the genus.

It grows to 30 m tall, with fragrant lance-shaped leaves, to 10 cm long. The yellow or yellowish-green flowers are arranged in small umbels, which has given rise to the generic name, meaning ‘little umbel’. The fruit is an oval berry, to 2.5 cm long, green at first, purple when mature.

The leaves have been used as a substitute for those of the true bay laurel (Laurus nobilis) in cooking. They were utilized by native tribes for a number of ailments, including headache, toothache, earache, stomach ache, colds, sore throat, and mucus in the lungs. A poultice of the leaves was used for rheumatism and neuralgia.

 

 

Flowering California bay laurel, Sequoia National Park. (Photos copyright © by Kaj Halberg)

 

 

California bay laurel with unripe fruits, Kings Canyon National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Shrubs

 

Arctostaphylos Manzanita, bearberry
A genus with about 70 species of mostly evergreen dwarf shrubs, shrubs, or small trees of the heath family (Ericaceae), widely distributed in the Northern Hemisphere, from the Arctic southwards to Central America, the Mediterranean, the Caucasus, and Mongolia.

Many American species are characterized by having smooth, orange or reddish bark, and stiff, twisting branches. Several species occur in the Sierras, many of them quite similar and often difficult to identify.

The generic name is derived from Ancient Greek arktos (‘bear’) and staphyle (‘a bunch of grapes’), alluding to the fruits, which form grape-like clusters in many species. They are much relished by bears.

The word manzanita is Spanish, a diminutive of manzana (‘apple’), referring to the fruits.

Red bearberry (A. uva-ursi), and a presumed hybrid between this species and hairy manzanita (A. columbiana), often called A. x media, are presented on the page Autumn.

 

Arctostaphylos viscida Whiteleaf manzanita
The ovate, pointed leaves of this shrub, growing to 5 m tall, are not really white, but whitish-green or dull green. It is widely distributed in California and Oregon. It is quite common in the lower montane zone of the Sierras.

In the past, the Miwok people of northern California made cider from the fruits.

The specific name is derived from the Latin viscidus (‘sticky’), alluding to the glandular leaves and fruits.

 

 

Reddish trunk of whiteleaf manzanita, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

Whiteleaf manzanita often produces an abundance of white or pinkish flowers. – Sequoia National Park. (Photos copyright © by Kaj Halberg)

 

 

Fallen flowers of whiteleaf manzanita, forming a carpet on the ground, Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Sambucus Elder
This genus of about 27 species of smaller trees, shrubs, or large herbs, is found mainly in temperate and subtropical areas of the Northern Hemisphere. On the Southern Hemisphere, they are restricted to parts of Australia and South America.

In the past, this genus and the genus Viburnum constituted the family Sambucaceae, but were then transferred to the honeysuckle family (Caprifoliaceae), and later to the moschatel family (Adoxaceae), which is today called Viburnaceae.

The generic name is derived from Ancient Greek sambuca, the name of an ancient instrument of Asian origin. Presumably, flutes were made from the branches. The name elder is derived from the Anglo-Saxon word aeld (’fire’). In those days, the hollow stems of elder were used to kindle a fire.

 

Sambucus cerulea Blue elder
This deciduous shrub, which can grow to a height of 9 m, was formerly known as Mexican elder (S. mexicana). In spring, it displays an abundance of butter-yellow flowers, followed by purplish-blue berries in autumn, which gave rise to its names.

It grows in a variety of habitats, including river valleys, woodlands, and exposed slopes with access to water, up to altitudes around 3,000 m. It has a very wide distribution, from British Columbia southwards through much of western United States to north-western Mexico, with scattered occurrence in Oklahoma and Texas.

Many indigenous tribes used the berries of this plant for food, and a number of herbal medicines were produced from its wood, bark, leaves, flowers, and roots. The wood was also used to make pipes and musical instruments, such as flutes and small whistles. A dye was obtained from the berries.

 

 

Blue elder, Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Herbs

 

Castilleja Indian paintbrush
This genus, also known as prairie-fire, contains about 200 species, most of which have brilliant red flowers and bracts, whereas a few are orange, yellow, or violet. The vast majority are native to the western parts of the Americas, from Alaska southwards to the Andes. One species, however, C. pallida, is distributed across Siberia, southwards to the Altai Mountains and westwards to the Kola Peninsula.

These plants of the broomrape family (Orobanchaceae) are parasitic, obtaining a large part of their nutrients from roots of other plants.

The flowers of some species are edible and were formerly consumed by various native tribes, but as these plants tend to absorb and concentrate selenium in their tissue, roots and green parts can be very toxic.

 

 

An unidentified Castilleja species from Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Diplacus aurantiacus Sticky monkey-flower, orange bush monkey-flower
This shrubby plant, formerly known as Mimulus aurantiacus, grows to 1.2 m tall, with sticky, narrow leaves, to 7 cm long. The flower colour may be yellow, orange, or red.

An attractive plant, distributed from south-western Oregon through much of California, just extending into Baja California.

Formerly, members of the Miwok and Pomo tribes used it medicinally for treatment of wounds, burns, diarrhoea, and eye trouble.

The specific name is Latin, meaning ‘orange-coloured’.

 

 

Yellow-flowered form of sticky monkey-flower, Sequoia National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Equisetum Horsetails
This genus of spore-bearing plants, containing about 18 species, is found in most parts of the world, except Australia and Antarctica.

These plants have a hollow, jointed, silica-containing stem with 3-40 longitudinal grooves. The leaves, often reduced to scales, are arranged in whorls at each joint. There are two distinct types, of which one has brown, fertile spring shoots and green, sterile summer shoots, which supply the photosynthesis. The other group has only green summer shoots, with terminal sporangies. They vary from dwarf plants, 20 cm tall, to giants up to 8 m tall.

The generic name is derived from the Latin equus (‘horse’), and seta, which has several meanings, including ‘rough’, ‘brush’, or ‘hair’. The latter word can refer to the rough, silica-containing stems of these plants, but together with equus, the word means ’horse hair’. With a bit of imagination, a bunch of drying stems do resemble a horsetail.

A number of horsetail species are described on the page Plants: Plants in folklore and poetry.

 

Equisetum hyemale Rough horsetail
This plant, also called scouring rush, has an enormous distribution, found in North America, Europe, and northern Asia, southwards to Tibet, Korea, and Japan. It has also been introduced to other countries, including South Africa and Australia, where it is regarded as an invasive.

It grows in wet areas in forests, often on slopes along streams. In Japan, it is cultivated in ponds in ornamental gardens.

Certain indigenous American tribes used a decoction of the stems for venereal diseases and as a diuretic.

The specific name is derived from the Latin hiems (‘winter’), and the suffix alis, thus ‘found in winter’, referring to the evergreen stems.

The name scouring rush stems from its former usage as a scouring remedy, and as sandpaper, due to its high content of silica.

 

 

This picture from Kings Canyon National Park shows the American subspecies of rough horsetail, ssp. affine, which is often yellow above each sheath. (Photo copyright © by Kaj Halberg)

 

 

 

Kumlienia False buttercups
A small genus with two species, K. hystricula (described below), and K. cooleyae, which is distributed along the coast from Alaska southwards to Washington. These plants were formerly included in the genus Ranunculus.

The generic name honours Swedish-born American naturalist Thure Ludwig Theodor Kumlien (1819-1888), who contributed much to the knowledge of the natural history of Wisconsin. He taught botany and zoology, as well as foreign languages, at the Albion Academy.

Some authorities, including Kew Gardens of London, do not accept this genus, but include it in Ranunculus.

 

Kumlienia hystricula Waterfall false buttercup
This herb was formerly regarded as a species of buttercup, named Ranunculus hystriculus. As its common name implies, it grows in wet places, such as meadows and along streams. It is endemic to the Sierras, where it is restricted to coniferous forests.

The specific name is derived from hystrix, the classical Greek word for porcupine. The connection to this plant is hard to see.

 

 

Waterfall false buttercup, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Letharia Wolf lichens
Wolf lichens, comprising at least 6 species in the family Parmeliaceae, are widely distributed in North America, Siberia, and Europe. At least 2 species, the common wolf lichen (L. vulpina) and the brown-eyed wolf lichen (L. columbiana), occur in the Sierras.

Native peoples of California utilized these plants for a variety of purposes. Arrow poison was produced from their toxic foliage. Klamath people would soak porcupine quills in an extract of the plants, which dyed them yellow. They were then woven into baskets to make patterns. The Okanagan-Colville tribe used these lichens medicinally, internally for stomach problems, externally for wounds.

The generic name is Latin, meaning ‘deadly’, alluding to the toxicity of these plants. In the old days, Norwegians used common wolf lichen to kill wolves and foxes. The toxic foliage was mixed with crushed glass, and then stuffed into a carcass on frozen ground. The specific name vulpina refers to foxes, in Latin Vulpes.

 

 

A species of wolf lichen, forming concentric rings on a trunk of white fir (see above), Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

Wolf lichen, growing on trunks of tamarack pines (see below), Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Madia 
A genus of 11 species of aromatic and pretty herbs of the aster family (Asteraceae), distributed in western North America and south-western South America.

The generic name is derived from madi, a native Chilean name of the coast madia (M. sativa). Their foliage exudes a fragrant oil, hence the common name tarweed.

 

Madia elegans Common madia
This plant, distributed from Washington southwards to central California and Nevada, is divided into a number of subspecies, whose flower colour varies from solid lemon-yellow to lemon-yellow with white or maroon centre. The ray florets curl up during the daytime, opening late in the afternoon and staying open all night until mid-morning.

The small nut-like fruits (achenes) were previously eaten raw by several indigenous tribes, or ground into flour and baked.

 

 

Common madia, ssp. vernalis, Sequoia National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Primula Primrose, cowslip, oxlip
This large genus, with about 500 species of herbs, is native to temperate areas of the Northern Hemisphere and of southern South America, and also to tropical mountains in Ethiopia, Indonesia, and New Guinea.

The leaves of these plants are arranged in a basal rosette, and the flowers are usually in terminal umbels on a leafless stalk. The petals are usually fused, forming a tube with 5 terminal, spreading lobes.

A large number are very hardy, reflected in the generic name, which is a diminutive of the Latin prima (’first’), referring to the early flowering of several species. The name primrose stems from the Latin prima rosa, meaning ‘the first rose’, although primroses are not even distantly related to roses.

According to some authorities, the name cowslip is a corruption of an Old English word, cuslyppe (‘cow dung’). This probably refers to the favoured habitat of the common cowslip (P. veris), namely dry slopes, grazed by cattle. Others claim that the word is a corruption of cow’s leek, derived from the Anglo-Saxon word leac (‘plant’).

It has been said that the name oxlip, which is a corruption of ox and slip, refers to the fact that oxlip, like cowslip, often grows on cattle grazing grounds. This may be a wrong presumption, as true oxlip (below) – at least in the northern part of its distribution area – mainly grows in forests.

The role of these plants in folklore and traditional medicine is described on the page Plants: Primroses. Shooting stars are a group of primroses, which were formerly placed in the genus Dodecatheon, derived from Ancient Greek dodeka (’twelve’) and theos (’god’), thus ‘twelve gods’. The Greeks applied this name to plants that they thought had acquired their medicinal properties from the twelve most prominent gods and goddesses. The genus was erected by Swedish naturalist Carl Linnaeus (1707-1778), supposedly in allusion to the inflorescence of these plants often consisting of 12 flowers.

This group, comprising 17 species, is widespread in North America, from north-western Mexico northwards through western United States and Canada to Alaska, with a single species extending into north-eastern Siberia. The flowers are pollinated by bees, which cling to the petals by beating their wings very fast, thus releasing the pollen.

The popular name shooting star refers to the flowers, in which the yellow stamens and the petals, which are bent backwards, converge to a common point, likened to a shooting star, in which the petals constitute the ‘tail’.

 

Primula jeffreyi Jeffrey’s shooting star
This species, previously named Dodecatheon jeffreyi, is distributed from Alaska and Montana southwards to California, growing in mountain meadows and along streams. An alternative name of it is Sierra shooting star.

The Nlaka’pamux people used the flowers as amulets and love charms.

It was named in honour of Scottish botanist John Jeffrey (1826-1854), who spent four years exploring and collecting plants in Washington, Oregon, and California, sending his specimens back to Scotland. In 1854, he disappeared while travelling from San Diego across the Colorado Desert, and was never seen again.

 

 

Jeffrey’s shooting star, Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

 

Piñon pine – juniper woodland (eastern dry slopes), c. 1500-2100 m

 

Trees

 

Juniperus Juniper
This genus of c. 60 species of trees and shrubs is distributed in almost the entire Northern Hemisphere. The cone is berry-like, with an outer fleshy, leathery layer. In Asia, foliage of junipers is often burned as incense at Buddhist and Hindu shrines.

There are various theories as to the origin of the generic name. Some authorities claim that it stems from the Latin iungere (’tie together’ or ’weave’), referring to the use of its branches in baskets and fences, whereas others maintain that it is derived from juvenis (’young’) and parere (’to produce’), referring to the fact that juniper bushes constantly are renewed by new shoots.

The words gin and genever are derived from Juniperus, attesting to the usage of juniper fruits in these beverages.

 

Juniperus osteosperma Utah juniper
This is a small tree, usually much stunted and only 3-6 m tall, which grows in very dry desert areas and on mountain slopes at moderate altitudes, from southern Montana and Idaho southwards to western New Mexico, westwards to the extreme eastern California. In the Sierras, it is restricted to the eastern slopes and the White Mountains.

The cones are berry-like, to 1.3 cm across, bluish-brown, with a whitish waxy bloom.

In former days, Native Americans used the bark for a variety of purposes, including beds, and ate the cones both fresh and in cakes. The gum was smeared on wounds as a protective covering. Tea made from the leaves was given to women to calm their contractions after giving birth. The Navajo would sweep their tracks with boughs from the tree, so that death would not follow them.

The specific name is derived from Ancient Greek osteon (‘bone’) and sperma (‘seed’), alluding to the hard seeds of this species.

 

 

Old Utah junipers often have strongly distorted trunks. These were observed in Canyon de Chelly National Monument, Arizona (top), and Inyo National Forest, White Mountains, California. (Photos copyright © by Kaj Halberg)

 

 

This stunted Utah juniper in Canyonlands National Park, Utah, has only a 10 cm wide strip of bark on the left side. (Photo copyright © by Kaj Halberg)

 

 

An unusually straight Utah juniper, Canyonlands National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Pinus monophylla Singleleaf piñon

 

The pine nuts [of Pinus monophylla] are delicious – large quantities are gathered every autumn. The tribes of the west flank of the range trade acorns for worms [caterpillars of a species of silkworm] and pine nuts. The squaws carry immense loads on their backs across the rough passes and down the range, making journeys of about forty or fifty miles each way.”

 

John Muir (1838-1914), Scottish-American writer and environmentalist, in his book My First Summer in the Sierra, 1911.

 

 

As its specific and common names imply, needles of this pine are not arranged in groups. It is the only pine in the world with solitary needles.

It is one among several pines of the piñon pine group, which are native to western United States and north-western Mexico, from southernmost Idaho, Utah, and south-western New Mexico westwards to eastern and southern California and Baja California.

This tree, which grows to 20 m tall, usually occurs at altitudes between 1,200 and 2,300 m. It is very common, forming open woodlands, often mixed with junipers.

As described in the quote above, the edible nuts were formerly an important source of food for indigenous peoples, and they are still widely harvested.

 

 

Pale morning light on woodland with forest of singleleaf piñon, Inyo National Forest, White Mountains. The Sierras are seen in the background. (Photo copyright © by Kaj Halberg)

 

 

Singleleaf piñon, Inyo National Forest. (Photo copyright © by Kaj Halberg)

 

 

Bark on an old singleleaf piñon, Inyo National Forest. (Photo copyright © by Kaj Halberg)

 

 

Cones and foliage of singleleaf piñon, Joshua Tree National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Upper montane forest, c. 2100-2700 m

 

Trees

 

Pinus contorta Lodgepole pine
One of the indicator species of this zone is the tamarack pine, or Sierra lodgepole pine, which is a subspecies, murrayana, of the widespread lodgepole pine. This species, divided into four subspecies, is very common in western North America, from sea level to subalpine montane forests.

Tamarack pine is widely distributed in mountains, from Washington southwards to northern Baja California, and thence eastwards to southern Nevada. The nominate contorta is found along the Pacific Coast from southern Alaska to north-western California. Another coastal subspecies is bolanderi, which is endemic to Mendocino County, north-western California. It is threatened by urban development. Finally, ssp. latifolia is found in the Rocky Mountains, from Yukon and Saskatchewan southwards to Colorado.

The specific name is derived from the Latin con (‘with’) and torqueo (‘twisted’), referring to the low, twisted trees, commonly found along the Pacific Coast. This is also reflected in one of the common names of this species, twisted pine.

Native peoples often used this species to build their lodges, hence the common name lodgepole pine.

 

 

One of the common names of the lodgepole pine is twisted pine, a suitable name for this specimen of tamarack pine in Yosemite National Park. (Photos copyright © by Kaj Halberg)

 

 

Cones and needles of tamarack pine, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Populus Poplar, aspen, cottonwood
This genus contains about 60 species of deciduous trees, some of which are indeed majestic, growing to 50 m tall, with a trunk diameter up to 2.5 m. They are native to the major part of the Northern Hemisphere, from subarctic areas southwards to Mexico, North Africa, Iran, the Himalaya, and China. Many species are planted elsewhere as ornamentals, or to produce timber.

The leaves are broad, mostly heart-shaped, with a long, slender stalk, which is flattened, making the leaf rustle in the wind. Inflorescences are pendent catkins, with numerous tiny flowers clustered around a central axis, each flower surrounded by papery bracts. The flowers are wind-pollinated, seeds fluffy, spread by the wind.

Several species display brilliant yellow foliage in the autumn, examples of which are shown on the page In praise of the colour yellow.

The generic name is the classical Latin word for poplars. The common name cottonwood alludes to the fluffy seeds. The name aspen is derived from Old English æspe, originally from Proto-Germanic aspe (‘shaking’), alluding to the fluttering leaves of European aspen (P. tremula).

 

Populus tremuloides Quaking aspen
A common tree in the upper montane zone. This species, which is called by many other names, including golden aspen and trembling poplar, is the most widely distributed tree in North America, found from Alaska southwards through western Canada and the United States to central Mexico, and in a broad belt across Canada and northern U.S. to Newfoundland and New England.

In autumn, the foliage of this iconic species adds vivid splashes of yellow to numerous montane areas in western North America.

 

 

Quaking aspens, displaying their brilliant autumn foliage on a mountain slope near Conway Summit. The lower two pictures show the snow-white trunk of this iconic species. (Photos copyright © by Kaj Halberg)

 

 

 

Shrubs

 

Chrysolepis Chinquapin
This small genus of the beech family (Fagaceae) contains only 2 species, one tree and one shrub, both restricted to the westernmost part of the United States, from Washington southwards through Oregon to California and Nevada.

The leaves of these plants have a dense layer of golden scales beneath, which gave rise to the generic name, derived from Ancient Greek khrysos (‘gold’) and lepis (‘scale’). The fruit is very spiny, a so-called cupule, containing three edible nuts, which were a common food of indigenous peoples, raw or roasted.

The name chinquapin also refers to the related genera Castanopsis and Castanea. This word is a corruption of an Algonquian word, chechinquamin or chincomen, possibly from xinkw (‘great’) and mini (‘fruit’).

 

Chrysolepis sempervirens Bush chinquapin
This small evergreen shrub, to 2 m tall, grows at high elevations, between 1,000 and 3,000 m. It is distributed from southern Oregon southwards to southern California, eastwards to the Sierras.

The specific name is Latin, meaning ‘evergreen’.

 

 

Fruiting bush chinquapin, Yosemite National Park. (Photo copyright © by Kaj Halberg)

 

 

 

Herbs

 

Helenium Sneezeweeds
A genus with about 35 species of the aster family (Asteraceae), distributed from Canada southwards to the Caribbean and Central America, and also found in southern South America.

Originally, Helenium was the name of elecampane (Inula helenium), which, it was said, Helen of Troy planted on the island of Pharos.

The common name was given in allusion to the ancient usage of dried leaves to make snuff, which was inhaled to help sneezing, thus ridding the body of evil spirits.

 

Helenium bigelovii Bigelow’s sneezeweed
This plant, found in California and Oregon, grows in wet areas, including meadows and along streams, at relatively high altitudes, from 1,000 to 3,000 m. Cultivated forms are widely grown as ornamentals.

The specific name was given in honour of surgeon and botanist John Milton Bigelow (1804-1878), who collected many new plant species on expeditions to south-western U.S. and northern Mexico.

 

 

Bigelow’s sneezeweed, Kings Canyon National Park. In the lower picture, a yellow-faced bumblebee (Bombus vosnesenkii) is feeding in the flowers. (Photos copyright © by Kaj Halberg)

 

 

 

Sarcodes sanguinea Snow plant
This parasitic member of the heath family (Ericaceae) derives its nutrients from underground fungi. It is distributed from the Cascade Range in Oregon through montane areas of California, including the Sierras, to northern Baja California, Mexico.

American botanist, chemist, and physician John Torrey (1796-1873) found the colour of this plant so striking that he named it Sarcodes sanguinea, from Ancient Greek sarkos (‘flesh’) and the Latin sanguis (‘blood’), thus ‘the blood-coloured, fleshy one’. The common name refers to the early flowering of this species, which often appears, when snow is still partly covering the ground.

In his book The Yosemite (1912), Scottish-American writer and environmentalist John Muir (1838-1914) writes the following about this species: The snow plant (…) is more admired by tourists than any other in California. It is red, fleshy, and watery and looks like a gigantic asparagus shoot. Soon after the snow is off the ground, it rises through the dead needles and humus in the pine and fir woods like a bright glowing pillar of fire. (…) It is said to grow up through the snow; on the contrary, it always waits until the ground is warm, though with other early flowers it is occasionally buried or half-buried for a day or two by spring storms. (…) Nevertheless, it is a singularly cold and unsympathetic plant. Everybody admires it as a wonderful curiosity, but nobody loves it as lilies, violets, roses, daisies are loved. Without fragrance, it stands beneath the pines and firs, lonely and silent, as if unacquainted with any other plant in the world; never moving in the wildest storms; rigid as if lifeless, though covered with beautiful rosy flowers.”

 

 

Snow plant, photographed in the Siskiyou Mountains, Oregon. (Photo copyright © by Kaj Halberg)

 

 

 

Subalpine zone, c. 2700-2900 m

 

Trees

 

Pinus albicaulis Whitebark pine
This pine is an indicator plant of the subalpine zone in the Sierras. It is native to montane areas, from British Columbia southwards through Washington, Idaho, Montana, and Wyoming, and with patchy occurrence in Oregon, Nevada, and California.

In favourable conditions, it may grow to almost 30 m tall, but at exposed locations it often becomes stunted and twisted.

The specific name is Latin, meaning ‘white-stemmed’ (i.e. ‘with a white trunk’).

 

 

An old, gnarled whitebark pine, standing on the rim in Crater Lake National Park, Oregon. Yellow lichens are growing on its exposed roots. Crater Lake evolved in the caldera of a collapsed ancient volcano, Mount Mazama. In the picture, Wizard Island is seen in the background. (Photo copyright © by Kaj Halberg)

 

 

 

Pinus longaeva Great Basin bristlecone pine
One day, towards the end of April 1992, I was hiking up a slope in Inyo National Forest, White Mountains – an eastern spur of the Sierras. In front of me were the most remarkable trees I have ever seen. At a distance, they appeared completely dead, with twisted, naked branches, stretching from a yellowish trunk towards the blue sky. But then – at closer quarters I noticed a narrow strip of bark on the side of the trunk, which pointed away from the direction of the prevailing wind. This strip of bark was leading up to one or two branches, densely covered in green needles, and from the tip of these branches, small cones were hanging down, their scales equipped with bristle-like appendages.

These peculiar trees are restricted to high-altitude areas in eastern California, and in Nevada and Utah. Apart from certain clones, including a creosote (Larrea tridentata) in the Mohave Desert, whose age is estimated at c. 9,400 years, this pine is the oldest living organism on Earth, a few of them being around 5,000 years old.

The specific name is Latin, meaning ‘ancient’.

Other photos, depicting these remarkable trees, are shown on the page Plants: Ancient and huge trees.

 

 

Ancient Great Basin bristlecone pines, Inyo National Forest, White Mountains. (Photos copyright © by Kaj Halberg)

 

 

 

(Uploaded December 2018)

 

(Latest update October 2024)