Algae and lichens

 

 

Cladonia floerkeana is a lichen with bright red sporangies, photographed on the island of Fanø, Denmark. (Photo copyright © by Kaj Halberg)

 

 

This Khmer ruin in Ta Prohm, Angkor, Cambodia, is covered in green algae. This fascinating area is described on the page Religion: Hinduism. (Photo copyright © by Kaj Halberg)
 

 

 

Orange algae, growing on a stone wall, Alishan National Forest, Taiwan. (Photo copyright © by Kaj Halberg)
 

 

 

This little girl is playing with green algae, possibly Ulva compressa (see below), Langeland, Denmark. (Photo copyright © by Kaj Halberg)
 

 

 

Old-man’s-beard lichens of the species Usnea longissima are waving in the wind, Ghunsa Valley, eastern Nepal. (Photo copyright © by Kaj Halberg)

 

 

Channelled wrack (Pelvetia canaliculata) is a brown alga, growing on rocky coasts, here at Loch Hourn, Scotland. (Photo copyright © by Kaj Halberg)

 

 

 

This kelp, oar-weed, tangle, devil’s-apron, sole-leather, or ribbon-weed – as various species are called – appeared to us a singularly marine and fabulous product, a fit invention for Neptune to adorn his car with, or a freak of Proteus.”

 

American writer Henry David Thoreau (1817-1862), in his book Cape Cod, first published 1865. – Neptune is the Roman sea god, whereas Proteus is his Greek counterpart.

 

 

 

But above all other kinds of moss*, which grow in the forests on trees, rocks and other places, the most famous one is Usnea, or Muscus cranii humani, meaning: ’That moss which grows on human skulls’, which, although rarely, is sometimes found on the skulls of miscreants, who have been beheaded, or otherwise done away with, and whose heads have been placed on a stake.”

 

Danish physician and herbalist Simon Paulli (1603-1680) about old-man’s-beard lichen (Usnea), in his book Flora Danica (1648). His life is described on the page People: Famous naturalists.

 

*In those days, botanists didn’t distinguish between mosses and lichens.

 

 

 

Algae is a term that covers a large and diverse group of organisms, of which the majority are aquatic and autotrophic (i.e. producing organic material through photosynthesis or through inorganic chemical reactions). Many forms lack most of the distinct cell and tissue types of typical land plants.

Algae range from one-celled microalgae like Chlorella, Prototheca, and Ochrophyta (diatoms) to multicellular forms, including the large seaweeds, of which the largest, the giant kelp (Macrocystis pyrifera, see below), may grow up to 50 m long. The most complex freshwater forms are the Charophyta, a division of green algae, which includes Spirogyra and Charales (stoneworts).

Cyanobacteria, popularly known as blue-green algae, mostly live in freshwater, producing energy through photosynthesis. Even though they are not really algae, they are included here due to their popular name. A few pictures of these organisms are shown at the bottom of the page.

Lichens are double organisms, consisting of a fungus and an alga or a cyanobacterium, living in a mutual symbiotic relationship. The alga or the cyanobacterium is attached to the mycelium of the fungus, which is again attached to the soil, or to stones or trees. Thus, the alga or cyanobacterium is protected from most enemies, and in return it provides the fungus with essential carbohydrates that the fungus cannot produce. The mycelium, however, is able to obtain certain nutrients from the soil or from the air, which benefit the alga or cyanobacterium.

Some species of marine algae form symbiotic relationships with other organisms than lichens or cyanobacteria, for instance corals or sea sponges.

 

Algae
Algae, comprising at least 50,000 species (probably many more than that), are divided into several groups, mostly distinguished by colour: green algae, stoneworts, red algae, brown algae, and yellow or yellow-green algae.

Green algae (Chlorophyta and Streptophyta) are a huge and diverse group, containing between 9,000 and 12,000 species. The name Chlorophyta is derived from Ancient Greek chloros (‘green’) and phyton (‘plant’), Streptophyta from Ancient Greek strepto (‘twisted’) and again phyton, where strepto alludes to the morphology of the sperm of some members.

Stoneworts (Charophyceae) are a class of green algae, comprising about 400 species worldwide, most of which grow in freshwater, a few in brackish water. These organisms may grow to 1.2 m long and are strongly branched, with leaves in whorls. They produce energy via photosynthesis. They are often abundant in clear lakes and ponds, but disappear with growing eutrophication, suffocated by increased production of plant plankton.

The name stonewort stems from a layer of calcium carbonate, which over time may be encrusted on these plants.

Red algae (Rhodophyta) are one of the oldest groups of algae, containing over 7,000 species, the vast majority consisting of marine algae, including many notable seaweeds, whereas they are rare in freshwater habitats. The name is derived from Ancient Greek rhodon (‘rose’) and phyton (‘plant’), thus ‘rose-coloured plant’.

Brown algae (Phaeophyceae) constitute a huge group of large algae, comprising between 1,500 and 2,000 species, including many species in colder waters of the Northern Hemisphere. In daily speech, they are called kelp or seaweeds. Some members of the group are consumed by people, especially in the Far East.

Most of the yellow or yellow-green algae (Xanthophyceae), also known as mustard algae, are yellowish, but may sometimes be bright orange. They mainly live in fresh water, but some are found in the sea, others on land, where they may cling to various surfaces, usually in humid places. They vary from single-celled flagellates to simple colonies and filamentous forms. Supposedly, their nearest relatives are the brown algae.

 

Fucaceae
A small family of brown algae with 6 genera and about 23 species.

 

Ascophyllum nodosum Knotted wrack
This species, also known as egg wrack, isthe only species in the genus. The fronds, to 2 m long, are olive-green or olive-brown, without midrib. They are inflated at regular intervals, forming large, egg-shaped air bladders.

It is restricted to the northern Atlantic Ocean, from Svalbard southwards to Portugal, in eastern Greenland and southwards to the north-eastern coasts of North America.

The name is derived from Ancient Greek askos (‘bag’ or ‘bladder’) and phyllon (‘leaf’), and from the Latin nodosus (‘knotted’), from nodus (‘joint’), referring to the air bladders on the fronds.

 

 

Knotted wrack, Loch Hourn, Scotland. (Photo copyright © by Kaj Halberg)

 

 

 

Fucus vesiculosus Bladder wrack
This species is found on the Atlantic shores of Europe, eastwards to northern Russia and the Baltic Sea, in Morocco, the Canary Islands, Madeira, and the Azores, and also from Greenland and eastern Canada southwards to North Carolina.

Attached to stones or rocks, the branched fronds of this plant may grow to 90 cm long and 2.5 cm wide. Along both sides of the prominent midrib are a number of almost globular air bladders.

Since 1811, this species has been a source of iodine, utilized to treat goitre, a swelling of the thyroid gland, caused by lack of iodine.

The generic name stems from Ancient Greek phukos, a term used for seaweeds. The specific name is derived from the Latin vesicula (‘small bladder’), and the suffix osus (‘full of’).

 

 

Bladder wrack, Loch Hourn, Scotland. (Photo copyright © by Kaj Halberg)

 

 

 

Washed-up bladder wrack and other brown algae are often used by breeding birds as nesting material.

 

 

Incubating common eider females (Somateria mollissima), well camouflaged among washed-up bladder wrack, Mågeøerne, Funen, Denmark. (Photos copyright © by Kaj Halberg)
 

 

 

Where the great cormorant (Phalacrocorax carbo ssp. sinensis) breeds on the ground, it often builds tall nests, consisting of washed-up bladder wrack, which, when dry, makes a stable foundation for the nest. – Mågeøerne, Funen (top), and Svanegrund, eastern Jutland, Denmark. (Photos copyright © by Kaj Halberg)
 

 

 

Cormorants often bring green plants to their nest. – Svanegrund. (Photo copyright © by Kaj Halberg)
 

 

 

Cormorant chicks in a nest, Mågeøerne. (Photo copyright © by Kaj Halberg)

 

 

Nest of herring gull (Larus argentatus) among washed-up bladder wrack, containing two chicks and an egg, Svanegrund, eastern Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Pelvetia canaliculata Channelled wrack
This plant, the only member of the genus Pelvetia, grows in dense, branched tufts, to 15 cm long, and the fronds are deeply channeled on one side, an aid to avoid drying out at low tide. It is common on the Atlantic shores of Europe, from Iceland and Norway southwards to Spain and Portugal.

In Ireland, where this species is known as dúlamán, it was collected for food during famines. A local folk song, Dúlamán, describes a growing relationship between two people who collect the seaweed as a profession.

The generic name was given in honour of French botanist Auguste Pelvet (1821-1859). The specific name is derived from the Latin canaliculus (‘small channel’), and the suffix atus (‘having’).

 

 

Channelled wrack, growing on stony coasts, Dervaig, Isle of Mull, Inner Hebrides (top), and Loch Hourn, both in Scotland. (Photos copyright © by Kaj Halberg)

 

 

Channelled wrack with a washed-up green alga, possibly a species of Chaetomorpha, Loch Hourn. (Photo copyright © by Kaj Halberg)

 

 

 

Laminariaceae
A family of brown algae with 11 genera and about 64 species.

The name is derived from the Latin lamina (a thin plate or layer) and the suffix arius (‘resembling’), referring to the large and flat fronds of many of these algae.

 

Macrocystis pyrifera Giant kelp
This is the largest of all algae, to 50 m long, capable of growing up to 60 cm a day. It forms dense stands along the coast from south-eastern Alaska southwards to Baja California, and is also found along the shores of southern South America, South Africa, Australia, and New Zealand.

The name is derived from Ancient Greek makros (‘long’ or ‘large’) and kystis (‘bladder’ or ‘sac’), and the Latin pyrus (‘pear’) and fero (‘to carry’), alluding to its large size, and to the numerous pear-shaped bladders that keep it floating.

 

 

This giant kelp, several stories high, is displayed in the Monterey Aquarium, California. Other pictures from this gorgeous aquarium may be seen on the pages In praise of the colours red and orange, and In praise of the colour blue. (Photo copyright © by Kaj Halberg)

 

 

This specimen was washed ashore on Melkbosstrand, near Cape Town, South Africa, and is now partly covered by sand. Table Mountain is seen in the background. (Photo copyright © by Kaj Halberg)

 

 

Steffen Madsen with giant kelp, washed ashore near Algeciras, southern Spain. (Photo copyright © by Kaj Halberg)

 

 

 

Nereocystis luetkeana Bull kelp
This plant forms thick stands, comprising an important part of the kelp forests along the Pacific Coast of North America, from the Aleutian Islands southwards to southern California. It is attached to rocks and has a cylindric stem to 36 m long, terminating in a dense cluster of fronds, to 10 m long and 15 cm wide.

The generic name is derived from Ancient Greek Nereis (originally a sea god, but in Latin meaning a mermaid), and kystis (‘bladder’), thus ‘mermaid’s bladder’.

The specific name was applied in honour of German-Russian navigator, geographer, and Arctic explorer Friedrich Benjamin Graf von Lütke (1797-1882), in Russia known as Fyodor Petrovich Litke.

 

 

Bull kelp, photographed at various locations in California. (Photos copyright © by Kaj Halberg)

 

 

Washed-up bull kelp creates patterns on a stony beach, Salt Point State Park, California. (Photo copyright © by Kaj Halberg)

 

 

Snowy egret (Egretta thula), walking on a ‘carpet’ of bull kelp, Point Lobos State Park, California. (Photo copyright © by Kaj Halberg)

 

 

 

Postelsia palmaeformis Sea palm
This plant, named for its superficial resemblance to palm trees, is the only member of the genus. It is distributed along the west coast of North America, growing on rocky shores with constant waves. It is one of the few brown algae that can survive and remain erect on land. In fact, it spends most of its life exposed to the air.

The generic name was given in honour of German-Estonian geologist and artist Alexander Philipov Postels (1801-1871).

 

 

Sea palm, growing on a rock, Salt Point State Park, California. (Photo copyright © by Kaj Halberg)

 

 

 

Trentepohliaceae
This family of terrestrial green algae contains about 5 genera with c. 100 species, known for their bright orange or red filaments, caused by high concentrations of carotenoid pigments, which cover their chlorophyll. They are often found on trees or wet rocks, living in symbiotic relationship with lichens. Members are found almost worldwide.

 

Trentepohlia aurea
This gorgeous alga is distributed almost worldwide, being most common in tropical and subtropical regions. It grows on a variety of substrates, including rocks, old walls, tree bark, rotting wood, mosses, and wet soil.

The generic name was given in honour of German clergyman and botanist Johann Friedrich Trentepohl (1748-1806). The specific name is Latin, meaning ‘golden’.

 

 

Trentepohlia aurea, growing on a rock, Point Reyes National Seashore, California. (Photo copyright © by Kaj Halberg)

 

 

In these pictures, Trentepohlia aurea grows on old Monterey cypresses (Cupressus macrocarpa) in Point Lobos State Park, California. This tree is described on the page Plants: Ancient and huge trees. (Photos copyright © by Kaj Halberg)

 

 

 

Ulvaceae
The vast majority of members of this family of green algae, comprising 9 genera with 120-150 species, live in the sea, with only 3 species growing in freshwater.

 

Ulva armoricana
This species was described as late as the mid 1990s from Brittany, north-western France. It flourishes in shallow waters, which are polluted by nitrates and phosphates from farming areas, forming large, floating masses, which wash up on beaches. Frequent blooming occurs in northern and western Brittany, and as the seaweed decays it releases large amounts of foul-smelling hydrogen sulfide gas, which is highly toxic and may affect people’s health in areas of high concentration.

The name Ulva can be traced back to a Celtic word for marsh plants, and it became a classical Latin term for sedges (Carex). Why Swedish naturalist Carl Linnaeus (1707-1778) applied the name to these sea plants is not known. – His life is described on the page People: Famous naturalists.

The specific name is derived from Armorica, the classical Latin name for the coastal region of Brittany. This word can be traced back to Celtic are-mori (‘by the sea’).

 

 

Ulva armoricana, snails, and limpets, St. Servan, near St. Malo, Brittany. (Photo copyright © by Kaj Halberg)

 

 

 

Ulva compressa
This species is often called thread weed, as its fronds are very narrow, up to 20 cm long. It is found almost worldwide, often growing on coastal rocks.

The specific name refers to the fronds.

 

 

At the small coastal town of Laomei, near the northernmost point of Taiwan, is an area of flat coastal rocks of volcanic origin, the major part of which are covered in Ulva compressa. (Photos copyright © by Kaj Halberg)

 

 

Continued erosion by waves at Laomei has created numerous channels in the rocks, some so narrow that when a large wave enters them, the water is pressed upwards like a fountain. (Photos copyright © by Kaj Halberg)

 

 

 

Ulva lactuca Sea lettuce
This species is very common around the world, growing on coastal rocks or attached to other sea organisms.

The specific name is Latin, meaning ‘lettuce’, applied in 1753 by Linnaeus (see above) due to its striking resemblance to garden lettuce (Lactuca sativa).

 

 

Sea lettuce, growing on coastal rocks, Jialeshuei, Kenting National Park, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Unidentified Ulva species, growing on stones, St. Servan, near St. Malo (top), and Mont St. Michel, both Brittany. (Photos copyright © by Kaj Halberg)

 

 

An unidentified Ulva species, possibly U. compressa, on a tidal flat during low tide, attached to the shell of a European cockle (Cardium edule), Fanø, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Unidentified algae

 

During a research trip in 1741 to the island of Öland, Sweden, Swedish naturalist Carl Linnaeus (1707-1778) paid a visit to a coastal area with flat, horizontal limestone rocks in the north-eastern part of the island. He named the place Neptuni Åkrar (‘Neptun’s Fields’) after the Roman god of the sea, comparing the long, flat rows of rock with the long, narrow fields of the 1700s.

Linnaeus is presented on the page People: Famous naturalists.

 

 

Limestone rocks with yellowish algae, Neptuni Åkrar, Öland. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae grow on these sculptures, which depict guardians of the Hindu Holy Water Temple in Ubud, Bali, Indonesia: a tiger and an ogre-like creature. (Photos copyright © by Kaj Halberg)

 

 

 

The former bulwark beneath the fort in St. Malo, Brittany, has become overgrown with green algae. (Photos copyright © by Kaj Halberg)

 

 

 

‘Soup’ of filamentous red algae, Cape of Good Hope, South Africa. (Photo copyright © by Kaj Halberg)

 

 

 

Young man with a lush growth of stoneworts (Charophyceae), eastern Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Drainage canal with green algae and fallen leaves of trees, including Madagascar almond (Terminalia mantaly), Taichung, Taiwan. This tree is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

 

Khmer ruins, covered in green algae, Ta Prohm, Angkor, Cambodia. Numerous pictures from this impressive area are shown on the pages Religion: Hinduism, and Decay. (Photo copyright © by Kaj Halberg)

 

 

Rubble from a Khmer temple, covered in green algae, Ta Prohm. (Photo copyright © by Kaj Halberg)

 

 

This Khmer relief at Bayon, Angkor Thom, depicting a Hindu goddess or a noble lady, is partly covered by orange algae. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae on a greenhouse, Funen, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Light and shadows on a wall along a canal, overgrown by green algae, Taichung, Taiwan. (Foto copyright © by Kaj Halberg)

 

 

 

These stairs in a Hindu temple near Ubud, Bali, Indonesia, are covered in green algae. A Chinese hibiscus (Hibiscus rosa-sinensis) has been left on the stairs as an offering. This plant is described on the page Plants: Mallows and allies. (Photo copyright © by Kaj Halberg)

 

 

 

Edible frog (Pelophylax esculentus) among lush vegetation of green algae and water-crowfoot (Ranunculus), Zealand, Denmark. The edible frog is described on the page Animals: Animal portraits. (Photo copyright © by Kaj Halberg)

 

 

 

Orange-red algae, grey lichens, and green mosses on a rock, Umbal Valley, Virgen Valley, Tyrol, Austria. (Photo copyright © by Kaj Halberg)

 

 

 

Pond with green algae, Bumpass Hell, Lassen Volcanic National Park, Cascade Range, California. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae on wrecked boats, Le Fret, Crozon, Brittany. (Photos copyright © by Kaj Halberg)

 

 

 

Brown algae, caught in grass on a beach, Kosa Ruskaya Koshka, Chukotka Peninsula, eastern Siberia. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae, growing on a surface root of a shoemaker tree(Hieronyma alchorneoides), Reserva Nacional Hacienda Baru, Costa Rica. This large forest tree, to 50 m tall, of the family Phyllanthaceae, is found from Mexico southwards through Central America to Brazil, Bolivia, and Peru, and also in the Caribbean. (Photo copyright © by Kaj Halberg)

 

 

 

Orange algae, growing on the walls of a school building, Alishan, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae, growing on road signs, Funen, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae among rice plants, Taichung, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

In the humid and mineral-rich air, which surrounds the Wai-O-Tapu Thermal Area, New Zealand, numerous trees and branches are covered by orange-yellow algae. (Photos copyright © by Kaj Halberg)

 

 

 

Coastal rocks, covered in green algae, Wanli, northern Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

In the Hindu legend TheChurning of the Milk Ocean, from theBhagavata-Purana, it is related that the gods had become weakened and had been usurped by the asuras (demons).

The gods appealed to the supreme god Vishnu for help, and he suggested that they should regain their power by drinking the miraculous amrita, the nectar of immortality, which they could obtain by churning the cosmic milk ocean, thus bringing the jar with amrita to the surface.

However, Vishnu advised the other gods to treat the asuras diplomatically by suggesting them to jointly churn the ocean. When the amrita was brought to the surface, Vishnu would ensure that the gods got hold of it.

The asuras readily agreed to participate in the churning. To perform this stupendous task, the gods and the asuras uprooted the mountain Mandara, placed it upside down in the ocean, and coiled the giant, many-headed naga (serpent) Vasuki around it. By pulling alternately at each end of Vasuki, the mountain would act as a gigantic churn, thus bringing the amrita to the surface.

The mountain, however, began sinking into the ocean floor, causing Vishnu to assume the shape of a giant, named Kurma, half man, half turtle. He then dove to the bottom of the sea, where he placed Mandara on his back, thus preventing the mountain from sinking.

Finally, the jar with amrita surfaced, whereupon a fierce battle between the gods and the asuras ensued, the latter grabbing the jar and running away with it.

Again, the gods appealed to Vishnu, who assumed the form of a new avatar, Mohini, a beautiful goddess, who seduced the asuras and managed to get hold of the jar of amrita, thus preventing evil from becoming eternal, and preserving the good.

 

 

Due to the role of Kurma in the legend The Churning of the Milk Ocean, turtles are sacred to Hindus. These sculptures, overgrown by green algae, were encountered in a temple in Ubud, Bali, Indonesia. (Photos copyright © by Kaj Halberg)

 

 

 

Red algae in sea water along a coast, Ottenby, Öland, Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae on the surface of a stagnant river, Wufong, Taichung, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae, covering the surface of a forest pond, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

The natural range of the Monterey pine (Pinus radiata) is limited to a few locations in Santa Cruz and San Luis Obispo Counties, central California, and two islands, Guadalupe and Cedros, off the west coast of northern Baja California, Mexico. In its natural range, it is seriously threatened by an introduced species of fungus, the pitch canker (Fusarium circinatum).

Elsewhere, however, this tree has been planted extensively, especially in Australia, New Zealand, Spain, Argentina, Chile, Uruguay, Kenya, South Africa, and the island of Tristan da Cunha.

 

 

The trunk of this Monterey pine in Wai-O-Tapu Thermal Area, New Zealand, is partly covered by an orange-yellow alga. (Photo copyright © by Kaj Halberg)

 

 

 

Red algae, living on the surface of ice, covering Reflection Lake, Mount Rainier, Washington, United States (top), and on snow, Sognefjellet, Norway. (Photos copyright © by Kaj Halberg)

 

 

 

Patterns in green algae, growing in a polluted stream, Taichung, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

 

These African oystercatchers (Haematopus moquini) blend in very well with dark algae, growing on coastal rocks, Muizenberg, near Cape Town, South Africa. This bird is described on the page Animals – Birds: Birds in Africa. (Photo copyright © by Kaj Halberg)

 

 

 

Sculpture, depicting a Hindu goddess, covered in green algae and cobwebs, Kintamani, Bali, Indonesia. (Photo copyright © by Kaj Halberg)

 

 

 

Car wreck with green algae, lichens, and moss, Blekinge, Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Washed-up red algae, Vangså Beach, Thy, Denmark (upper 2), and Mols, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

This shrivelled moon jellyfish (Aurelia aurita) got stuck on a washed-up plank, covered in green algae, Horsens Fjord, Denmark. It somewhat resembles a sad alien. (Photo copyright © by Kaj Halberg)

 

 

 

Orange alge on the trunk of a spruce, Thy, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

The trunk of this Indian horsechestnut (Aesculus indica) in Great Himalayan National Park, Himachal Pradesh, northern India, is partly covered by a bright yellow alga. This tree is described on the page Plants: Himalayan flora 3. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae and ivy (Hedera helix), growing on a fence, Funen, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

The trunk of this dead sycamore maple (Acer pseudoplatanus) is covered by green algae, Nature Reserve Vorsø, Horsens Fjord, Denmark. This tree is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae, growing on eroded coastal rocks, Montaña de Oro State Park, California. (Photo copyright © by Kaj Halberg)

 

 

 

An unidentified species of brown alga, Hermanus, South Africa. (Photo copyright © by Kaj Halberg)

 

 

 

This house wall in Alishan National Forest, Taiwan, has acquired patina from an orange alga, growing on it. (Photo copyright © by Kaj Halberg)

 

 

 

This pond in the Baishuei Terraces in the Yunnan Province, south-western China, is brimming with yellow algae. These terraces consist of snow-white calcium bicarbonate, which has been deposited by water, seeping down a slope. Over the millennia, terraces have been formed, which are sacred to the local Nakhi-people, described on the page People: Chinese minorities. (Photo copyright © by Kaj Halberg)

 

 

 

‘Soup’ of green algae, with bits of foam plastic, Vangså River, Thy, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

The outline of orange and green algae on a wet rock in a park in Taichung, Taiwan, resembles a rather sleepy dinosaur. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae, growing on hairy bracket (Trametes hirsuta), Funen, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

Dried-up layer of algae in a pond in a stone quarry, Djupvik, Öland, Sweden. (Photos copyright © by Kaj Halberg)

 

 

 

Green algae on the surface of a pond, Nature Reserve Vorsø, Horsens Fjord, Denmark. (Photos copyright © by Kaj Halberg)

 

 

 

In the humid air in the Mangapohue River Gorge, New Zealand, these yellow algae have managed to fasten themselves on a wire netting. (Photo copyright © by Kaj Halberg)

 

 

 

Green algae in a pond, Stege, Møn, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Washed-up brown algae have attained a reddish colour, Kapelludden, Öland, Sweden. The ruin in the background is Sankta Brita’s Kapell (‘Saint Birgitta’s Chapel’). The story behind this chapel is related on the page Religion: Christianity. (Photo copyright © by Kaj Halberg)

 

 

 

Algae in a pond, north of Aarhus, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Lichens
About 20,000 species of lichens are known, appearing in numerous forms and colours. They may be crust-like, firmly fixed to stones or the like, or they may be bushy, with plant-like foliage. In the old days, botanists did not distinguish between lichens and mosses, and still today the term moss is used for some groups, for instance reindeer moss and Iceland moss.

Lichens can grow on almost any surface, and they occur from sea level to high mountains. In the Himalaya, they have been found at elevations above 6000 m. They are very common on stones and tree bark, or they may grow on plants, fungi, or even other lichens. However, the vast majority are merely epiphytes, only a few being parasitic.

 

Cladoniaceae
Comprising 18 genera with about 560 species, this is one of the largest lichen families. Members are found on all continents, including Antarctica.

 

Cladonia Cup lichens and reindeer lichens
A diverse genus with about 230 species, found in most parts of the world. They can be divided into two groups, simple species with cup-shaped thallus, and species with bushy thallus, often called reindeer lichens.

Cup lichens are widely distributed in the Northern Hemisphere, found on the ground or on rocks. The thallus varies in shape from simple and pointed to cup-shaped and can be yellow, gray, or brown in colour.

Reindeer lichens cover vast areas in the northern tundra and taiga areas, and although they grow very slowly (3-5 mm per year), they constitute a very important food item for reindeer or caribou (Rangifer tarandus), moose or elk (Alces alces), and musk ox (Ovibos moschatus).

Cup lichens contain a gummy or starchy material, which, boiled with milk or syrup, has been used as a remedy for whooping cough and chest ailments.

In Scandinavia, reindeer lichens have been used in the manufacture of alcohol. In Denmark, in the 1700s, these lichens were harvested in great quantities and, mixed with heather (Calluna vulgaris) used as winter fodder for cattle and sheep.

The generic name is derived from Ancient Greek klados (‘branch’), thus ‘branched’ – an apt name for many members of the genus.

 

 

Dunes, clad in vegetation of reindeer lichen, presumably cream-coloured cup lichen (C. portentosa), and also crowberry (Empetrum nigrum) and heather, Hansted Wildlife Reserve, Thy, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cladonia floerkeana
This species, easily identified by its bright red sporangia, is widely distributed in the Northern Hemisphere.

It is known by several popular names, including gritty British soldiers, Bengal matches, and Devil’s matchsticks, all alluding to the red sporangia, situated at the end of a short stalk. In the old days, uniforms of British soldiers were bright red. Bengal matches were mostly produced in India, hence the name. They were larger than ordinary matches and burned with a larger, brigtly coloured flame, hence the name Devil’s matchsticks.

The specific name honours German botanist and lichenologist Heinrich Gustav Flörke (1764-1835), who initially studied theology, mathematics, and medicine, but in 1816 he was appointed professor of natural history at the University of Rostock, where he remained for the rest of his life. He specialized in lichens, primarily studying the genus Cladonia. For a number of years, he was an editor of Oekonomische Encyklopädie (‘Economic Encyclopedia’).

 

 

Cladonia floerkeana, central Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Cladonia stellaris Star-tipped reindeer lichen
This species is very common in subarctic and temperate areas of North America, Europe, and Asia, often covering huge areas in boreal forests. In Nepal, it has been reported from an elevation of 4,500 m.

The specific name is Latin, meaning ‘star-like’, referring to the radiating tips of the branches.

 

 

Star-tipped reindeer lichen, Lake Femunden, Hedmark, Norway. In the upper picture, leaves of common blueberry (Vaccinium myrtillus) are also present. (Photos copyright © by Kaj Halberg)

 

 

 

Parmeliaceae
This large, diverse family, containing about 71 genera with more than 2,700 species, is found in almost all corners of the world, from tropical rainforests to the polar regions.

 

Letharia Wolf lichens
Wolf lichens, comprising about 6 species, are widely distributed in North America, Siberia, and Europe.

The generic name is derived from the Latin letalis (‘deadly’), referring to the high toxicity of vulpinic acid, a chemical produced by the lichen. In the old days, the common wolf lichen (L. vulpina) was utilized by the Norwegians to kill wolves and foxes. Its foliage was mixed with crushed glass, and then stuffed into a carcass on frozen ground. The specific name refers to foxes, in Latin Vulpes.

Common wolf lichen and brown-eyed wolf lichen (L. columbiana) are both found in California, where many native peoples formerly utilized them for a variety of purposes. Their foliage was used to produce arrow poison. Klamath people would soak porcupine quills in an extract of the plants, which dyed them yellow. They were then woven into baskets to make patterns. The Okanagan-Colville tribe used these lichens medicinally, internally for stomach problems, externally for wounds.

 

 

This picture shows an unidentified species of wolf lichen, growing in concentric rings around the trunk of a white fir (Abies concolor), Sequoia National Park, California. The white fir is described on the page Plants: Plants of Sierra Nevada. (Photo copyright © by Kaj Halberg)

 

 

An unidentified species of wolf lichen, growing on the trunk of lodgepole pine (Pinus contorta) (upper 2), on the trunk of a common manzanita (Arctostaphylos manzanita, centre), and on a dead twig on a conifer, all in Yosemite National Park, California. The lodgepole pine is described on the page Plants: Plants of Sierra Nevada, manzanitas on the page In praise of the colours red and orange. (Photos copyright © by Kaj Halberg)

 

 

In this picture, wolf lichen is growing on a dead, fallen branch, Summit Lake, Lassen Volcanic National Park, Cascade Range, California. (Photo copyright © by Kaj Halberg)

 

 

 

Usnea Old-man’s-beard lichens
A large, almost worldwide genus with about 600 species, most of which are greyish-green and grow on trees. They are ubiquitous in wetter areas around the world, where they often drape trees, hanging down from twigs and branches, waving in the wind.

The medicinal usage of old-man’s-beard lichens is described on the page Plants: Plants in folklore and poetry.

The generic name derives from Arabic ušna (‘moss’). In former days, botanists didn’t distinguish between mosses and lichens.

 

 

Several Usnea species grow in montane areas of East Africa. These pictures are from Ruwenzori National Park, Zaire (top), Mount Meru, Arusha National Park, Tanzania (centre), and Aberdare National Park, Kenya. (Photos copyright © by Kaj Halberg)

 

 

In these pictures, Usnea lichens grow on an old wooden fence, Kapsait, Cherangani Hills, Kenya. (Photos copyright © by Kaj Halberg)

 

 

An unidentified species of Usnea, attached to the bark of an old pehuen (Araucaria araucana), Conguillio National Park, Chile. (Photos copyright © by Kaj Halberg)

 

 

 

Usnea longissima
This species is widespread in Eurasia and western North America, often draping branches and twigs of trees, hanging down and waving in the wind.

The specific name is Latin, meaning ‘very long’.

 

 

Usnea longissima, Buttle Lake, Vancouver Island, Canada (top), and Hoh Forest, Olympic National Park, Washington, United States. (Photos copyright © by Kaj Halberg)

 

 

Usnea longissima is ubiquitous in wetter areas of the Himalaya, in this picture draping the branches of a Himalayan birch (Betula utilis), near Pungi Tenga, Khumbu, eastern Nepal. This tree is described on the page Plants: Himalayan flora 1. (Photos copyright © by Kaj Halberg)

 

 

Usnea longissima, draping the branches of a species of maple, Acer caudatum, Deboche, Khumbu, eastern Nepal. Some Himalayan maple species are described on the page Plants: Himalayan flora 3. (Photos copyright © by Kaj Halberg)

 

 

 

Xanthomaculina
A small genus with 3 species, restricted to deserts and other arid areas in southern Africa and Namibia. These peculiar lichens are not attached to stones or trees as most other lichens, but are blown about by the wind, getting nourishments from organic debris and from micro-organisms in the air.

The first part of the generic name is derived from Ancient Greek xanthos (‘yellow’), referring to the yellowish upper surface of these lichens. The last part is Latin, meaning ‘spotted’. It is not clear what it refers to.

 

Xanthomaculina convoluta
This species is endemic to the Namib Desert in western Namibia.

The specific name is derived from the Latin convolutus (‘to roll up’ or ‘to entwine’), referring to the thallus, which in dry periods curls into tight, twisted scrolls, which unroll when conditions again become humid.

 

 

Dried, rolled-up Xanthomaculina convoluta, Namib Desert. (Photo copyright © by Kaj Halberg)

 

 

 

Peltigeraceae
A family with about 15 genera and 600 species, for the major part occurring in mountains in subtropical and tropical regions.

 

Lobaria Lung lichens, lungworts
This genus of foliose lichens, comprising about 60 species, is widespread around the world. They are unusual by having a three-part symbiosis, containing a fungus, an alga, and a cyanobacterium, the latter being able to fix nitrogen.

Previously, these organisms constituted a separate family, Lobariaceae, but have since been moved to Peltigeraceae.

The generic name is Latin, meaning ‘lobed’, ultimately derived from Ancient Greek lobos (‘a lobe of the liver’), alluding to the lobed thallus. The common names likewise refer to the surface of the thallus, which resembles that of a lung.

 

 

A species of lung lichen, possibly Lobaria himalayensis, Barun Valley, Makalu-Barun National Park, eastern Nepal. (Photo copyright © by Kaj Halberg)

 

 

 

Lobaria pulmonaria Common lung lichen
This species is widely distributed in Eurasia, Africa, and North America, primarily growing in humid areas.

The specific name is derived from the Latin pulmo (‘lung’), like the generic and common names alluding to the surface of the thallus, which resembles that of a lung.

 

 

Common lung lichen, covering a rock, Pawtuckaway State Park, New Hampshire, United States. (Photos copyright © by Kaj Halberg)

 

 

Common lung lichen with apothecia (fruiting bodies), Acadia National Park, Maine, United States. (Photo copyright © by Kaj Halberg)

 

 

 

Teloschistaceae
This huge family with about 115 genera and 650-1000 species occurs in the major part of the globe, with the largest diversity in temperate regions.

The name is derived from Ancient Greek telos (‘end’) and schizein (‘to split’), alluding to the finely divided, split ends of the thallus branches, characteristic of the type genus Teloschistes.

 

Xanthoria Orange lichens, golden shields
A cosmopolitical genus, whose members are common on tree bark, walls, and rocks. They may be identified by the distinctive sporocarps (fruiting bodies), popularly called ‘fairy cups’.

The generic name is derived from Ancient Greek xanthos (‘yellow’) and the suffix oria (‘close to’), referring to the orange-yellow colour of these lichens.

 

 

An unidentified species of orange lichen, possibly Xanthoria elegans, growing on a rock, Grossglockner, Austria. (Photos copyright © by Kaj Halberg)

 

 

 

Xanthoria aureola Seaside orange lichen
This species grows on coastal rocks. As opposed to the common orange lichen (below), it produces very few sporocarps.

The specific name is Latin, meaning ‘golden’, of course alluding to its colour.

 

 

Seaside orange lichen, growing on cracked coastal rocks, near Simrishamn, Skåne, Sweden. The green plants are thrift (Armeria maritima), described on the page Plants: Urban plant life. (Photo copyright © by Kaj Halberg)

 

 

Seaside orange lichen, forming patterns on a coastal rock, Bornholm, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Seaside orange lichen, growing on the concrete wall of a German bunker from World War II, Fanø, Denmark. (Photo copyright © by Kaj Halberg)

 

 

 

Xanthoria parietina Common orange lichen
This species, also called yellow scale, is distributed in most of Europe, Africa, Asia, Australia, and North America. It is widespread and common on rocks, large trees, roofs, walls, etc., as it is one of the few lichens, which is favoured by eutrophication.

The specific name, from the Latin paries (‘wall’), was applied by Swedish naturalist Carolus Linnaeus (1707-1778) in 1753. His life is described on the page People: Famous naturalists.

 

 

Common orange lichen and the fern wall-rue (Asplenium ruta-muraria), growing on an old wall in the city of Enkhuizen, West-Friesland, Holland. Wall-rue is described on the page Plants: Flora in the Alps and the Pyrenees. (Photo copyright © by Kaj Halberg)

 

 

This old tombstone on the island of Christiansø, Bornholm, Denmark, has been overgrown by common orange lichen. (Photo copyright © by Kaj Halberg)

 

 

Common orange lichen, growing on a branch of an old common oak (Quercus robur), Bornholm. Plenty of sporocarps are present. (Photos copyright © by Kaj Halberg)

 

 

Common orange lichen, forming concentric rings on a stone wall, Hammershus, Bornholm. (Photo copyright © by Kaj Halberg)

 

 

Common orange lichen and a grey lichen species on a clay jar, Torslunda Churchyard, Öland, Sweden. (Photo copyright © by Kaj Halberg)

 

 

 

Unidentified lichens

 

 

These coastal rocks at Cap Frehel, Brittany, are covered in greyish lichens. (Photo copyright © by Kaj Halberg)

 

 

Yellow lichens, growing on spines of the cactus species Eulychnia acida, Valle del Encanto, Ovalle, Chile. (Photos copyright © by Kaj Halberg)

 

 

In this picture, a yellow lichen is growing on the cactus species Copiapoa malletiana, Llanos de Challe National Park, Chile. Both cactus species are described on the page Plants: Cacti. (Photo copyright © by Kaj Halberg)

 

 

Trunk of a European larch (Larix decidua) with a pale green lichen, possibly an Usnea species, Valsaparence, Gran Paradiso National Park, Italy. Larch is described on the page Plants: Flora in the Alps and the Pyrenees. (Photo copyright © by Kaj Halberg)

 

 

Yellow lichens grow on the church Beit Giorgis, one of the famous rock churches in Lalibela, Ethiopia. Some of these churches are described on the page Travel episodes – Ethiopia 1996: Timkat – a Christian festival. (Photo copyright © by Kaj Halberg)

 

 

Orange and grey lichens on a rock, Col du Mt. Cenis, French Alps. (Photo copyright © by Kaj Halberg)

 

 

Stone wall with grey lichens, Frederiksø, Bornholm, Denmark. (Photo copyright © by Kaj Halberg)

 

 

This stone cross on a grave at Källa Gamla Kyrka (‘Källa Old Church’), Öland, Sweden, is covered by lichens. (Photo copyright © by Kaj Halberg)

 

 

Colourful orange and yellow lichens on a rock, Pinnacles National Park, California. (Photo copyright © by Kaj Halberg)

 

 

King’s Men, or Rollright Stones, a Neolithic stone circle near Chipping Norton, Oxfordshire, England. The stones are covered by lichens. (Photos copyright © by Kaj Halberg)

 

 

Lichens, forming patterns on an overturned stelae from the Khmer culture, Ta Prohm, Angkor Wat, Cambodia. This fascinating area is described on the pages Religion: Hinduism, and Decay. (Photos copyright © by Kaj Halberg)

 

 

Various lichens on rocks, Zolotoi Khrebet (‘Golden Ridge’), near Anadyr Airport, Chukotka, eastern Siberia. To the left a withering marsh Labrador-tea (Rhododendron tomentosum). (Photo copyright © by Kaj Halberg)

 

 

Ring-shaped lichens on a tree trunk, Sun-Moon Lake, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

Orange lichens on rocks, Sanetti Plateau, Bale Mountains National Park, Ethiopia. (Photo copyright © by Kaj Halberg)

 

 

Lichens grow on an oldfashioned fence, consisting of stacked spruce poles, Stensjö, north of Oscarshamn, Småland, Sweden. (Photos copyright © by Kaj Halberg)

 

 

Lichens and a species of stonecrop (Sedum), growing at an elevation of c. 3200 m, Thangshyap, Langtang National Park, Nepal. (Photo copyright © by Kaj Halberg)

 

 

Tree bark with lichens, Sanyi, Taiwan. (Photo copyright © by Kaj Halberg)

 

 

Lichens, growing on the trunks of old ash trees (Fraxinus excelsior), Funen, Denmark. Moss is also seen in the upper picture. Ash is described on the page Plants: Ancient and huge trees. (Photos copyright © by Kaj Halberg)

 

 

Stone slabs with carved Tibetan Buddhist mantras (so-called mani stones), erected by devout Buddhists, Langtang Village, Langtang National Park, Nepal. Over time, these slabs become covered in colourful lichens. (Photos copyright © by Kaj Halberg)

 

 

Old stone fence in morning light, overgrown by mosses and lichens, Drymen, Loch Lomond, Scotland. (Photo copyright © by Kaj Halberg)

 

 

Lichens on the trunk of an American holly (Ilex opaca), Fire Island, Long Island, New York State. This tree is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

A rock wall and a rock in Matobo National Park, Zimbabwe, are covered in yellow and reddish lichens. (Photos copyright © by Kaj Halberg)

 

 

Various lichens on a rock, Schirnaya Mountains, Chukotka, eastern Siberia. (Photo copyright © by Kaj Halberg)

 

 

White lichens grow profusely on the trunk of an old northern white-cedar (Thuja occidentalis), Eagle Lake, Acadia National Park, Maine, United States. This tree is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

Bark of an old variable oak (Quercus variabilis) with lichens, Wuling National Forest, Taiwan. This species is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

Yellow lichens on a rock, Ghora Tabela, Langtang National Park, Nepal. (Photo copyright © by Kaj Halberg)

 

 

Orange lichens on a rock, Zolotoi Khrebet (‘Golden Ridge’), near Anadyr Airport, Chukotka, eastern Siberia. (Photo copyright © by Kaj Halberg)

 

 

Lichens, growing on a fence pole with a ceramic insulator and rusted wire, Store Bredlund Plantation, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Various lichens, growing on the trunk of an old Lithocarpus tree, Sun-Moon Lake, Taiwan. This tree is described on the page Plants: Ancient and huge trees. (Photo copyright © by Kaj Halberg)

 

 

 

Cyanobacteria (Cyanophyceae)
These organisms, popularly known as blue-green algae, live in freshwater and obtain energy via photosynthesis. Their name stems from their colour, derived from Ancient Greek kyanos (‘blue’). Modern botanists do not regard these organisms as algae, but I include them here due to their popular name.

Proliferate reproduction of these ‘algae’ often takes place late in the summer. They sometimes occur in huge masses, dying ponds and lakes a characteristic bluish-green colour, which is often an indication of pollution with phosphates and other fertilizers to a level that may constitute a health hazard.

 

 

This female mallard (Anas platyrhynchos) is swimming about in a ‘soup’ of blue-green algae, Skanderborg Lake, Jutland, Denmark. The mallard is described on the page Animals – Animals in culture and religion: Anatidae. (Photo copyright © by Kaj Halberg)

 

 

Blue-green algae in Århus River, Jutland, Denmark. (Photo copyright © by Kaj Halberg)

 

 

Blue-green and green algae in an alkaline lake, Lake Bogoria, Kenya. (Photo copyright © by Kaj Halberg)

 

 

 

(Uploaded September 2021)

 

(Latest update July 2026)